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Wheat shorts, wheat middlings and feed flour

Datasheet

Description
Click on the "Nutritional aspects" tab for recommendations for ruminants, pigs, poultry, rabbits, horses, fish and crustaceans
Common names 

English: Wheat middlings; wheat shorts; wheat feed flour; wheat mill run; wheat millrun; wheat pollard / pollards ; wheat red dog. Other terms may or may not be generic: wheatfeed; wheat offals.

French: remoulage (remoulage bis, remoulage demi-blanc, remoulage blanc); farine basse de blé; farine fourragère (wheat feed flour); issues de meunerie (milling by-products, may include bran). German: Weizennachmehl (middlings, particularly starch-rich material); Weizenfuttermehl (wheat feed meal). Spanish: harinillas de trigo (middlings); salvado fino or afrechillo may refer regionally to finer wheat milling by-products. Italian: cruschello, farinetta, farinaccio, and tritello; collectively: cruscami. Portuguese: farelo fino de trigo (fine wheat milling by-products). Dutch: tarwegriess (wheat middlings). Swedish: vetefodermjöl, veteklifoder (wheat feed meal; middlings). Finnish: vehnärehujauho (wheat middlings); vehnärehujauho, alkiopitoinen (germ-rich wheat middlings). Polish: śruta / produkty uboczne przemiału pszenicy (wheat milling by-products) Ukrainian: пшеничне кормове борошно (wheat feed flour). Bulgarian: пшенично фуражно брашно (wheat feed flour). Russian: пшеничная мучка (wheat milling by-products). Czech: pšeničná krmná mouka (wheat feed flour; wheat milling by-products). Arabic: مخلفات مطاحن القمح (wheat milling by-products). Chinese: 小麦次粉 (wheat middlings); local names include 黑面, 黄粉, 下面 and 三等粉. Japanese: 小麦末粉 (komugi sueko; low-grade/last-run wheat flour fraction used as a feed material). Korean: 밀 미들링 (mil mideulling; transliterated name). Korean tariff terminology uses the broader category 밀기울 (wheat milling residues), which can also include bran and therefore is not a precise synonym. Vietnamese: cám mì (wheat milling offal/middlings in feed usage; local usage can overlap with bran). Thai: รำข้าวสาลี (wheat milling by-products, may include bran). Indonesian: hasil samping penggilingan gandum (wheat milling by-products). Hindi: गेहूँ की मिलिंग के उप-उत्पाद (wheat milling by-products). Tamil: கோதுமை அரவைத் துணைப்பொருட்கள் (wheat milling by-products). Swahili: pumba laini ya ngano (fine wheat pollard; attested trade wording in Tanzania). Afrikaans: koringvoer (wheat feed; broad feed designation).

Description 

Wheat middlings, wheat shorts, and wheat feed flour are by-products of the milling of the wheat grain (Triticum aestivum L. or other domestic Triticum species) for flour manufacture. In roller milling, wheat is conditioned with water before it is gradually broken between rolls and sifted; intermediate particles are repeatedly purified, reduced and sifted to recover additional flour. Bran, germ, low-grade flour and other fractions are separated at different points, then may be sold separately or blended into feed by-products. About 25-30% of the grain is thus made available as by-products, which have been traditionally used for livestock feeding. The selected milling technology, flour extraction target and mill-stream blending therefore determine the composition of the final product, notably its relative proportions of fibre and starch, driven respectively by the contents in bran and flour. Wheat middlings, shorts, and feed flour are less fibrous materials than wheat bran, as they include less envelopes and more floury endosperm particles (Blasi et al., 1998).

The definitions of those products depend on the milling traditions of their country of origin. As shown in the Common names section, many countries use a single generic name that may or may not include the bran. Cultures with a long tradition of wheat milling have names specific to each by-product that are then transposed into standards or regulations.

The American Association of Animal Feed Control Officials lists five products (AAFCO, 2002), sorted below by fibre content:

  • Wheat flour: it consists principally of wheat flour together with fine particles of wheat bran, wheat germ, and the offal from the "tail of the mill." It must contain not more than 1.5% crude fibre.
  • Wheat red dog: it consists of the offal from the "tail of the mill” together with some fine particles of wheat bran, wheat germ, and wheat flour. It must contain not more than 4% crude fibre.
  • Wheat shorts: it consists of fine particles of wheat bran, wheat germ, wheat flour, and the offal from the "tail of the mill". It must contain not more than 7% crude fibre.
  • Wheat middlings: it consists of fine particles of wheat bran, wheat shorts, wheat germ, wheat flour, and some of the offal from the "tail of the mill". It must contain not more than 9.5% crude fibre.
  • Wheat mill run: it consists of coarse wheat bran, fine particles of wheat bran, wheat shorts, wheat germ, wheat flour, and the offal from the "tail of the mill". It must contain not more than 9.5% crude fibre.

The Commission Regulation (EU) No 68/2013 lists two products (European Commission, 2022): 

  • Wheat middlings: Product of flour manufacture obtained from screened grains of wheat or dehusked spelt. It consists principally of particles of endosperm with fine fragments of the outer skins and some grain screenings.
  • Wheat feed: Product of flour or malting manufacture obtained from screened grains of wheat or dehusked spelt. It consists principally of fragments of the outer skins and of particles of grain from which less of the endosperm has been removed than in wheat bran.

Note that the EU-defined middlings are supposed to be more starchy than the EU-defined wheat feed, while the AAFCO-defined middlings are more fibrous than the feed flour, the red dog and the shorts. In French milling terminology, there are indeed several types of remoulages, from the coarse remoulage bis ("gray") to the finer remoulage blanc ("white", due to endosperm-derived material), with some products in between (remoulage demi-blanc, half-white). The term remoulage itself refers to a second milling step, and there are other names of milling by-products (recoupe, repasse, rebulet...). The French version of the Commission Regulation (EU) No 68/2013 translates Wheat middlings as Farine basse de blé and Wheat feed as Remoulage de blé, which is not really consistent and shows how confusing these terminologies can be. As a result, and despite attempts at formal definitions, the livestock industry tends use these names interchangeably (Blasi et al., 1998).

This family of wheat milling by-products should be regarded as a continuum ranging from coarse bran to food-grade flour. The products differ mainly in their fibre and starch contents, and sometimes in colour, with no clear boundaries between them other than those defined arbitrarily by thresholds for crude fibre (as in the AAFCO definition) or starch (as in the tables accompanying this datasheet). These by-products are valuable feed ingredients, rich in energy and nutrients, and suitable for all classes of livestock. However, their composition is highly variable, and it is therefore advisable to have representative samples analysed before use.

Distribution 

Wheat milling by-products are generated wherever wheat is milled for flour. The main wheat producers in 2025 were the EU, China, India, the Russian Federation, the USA, and Canada, who accounted for 70% of a world production of 843 million tons, with 540 million tons being used for food (FAO, 2026). Considering a flour extraction rate of 70-75% (Blasi et al., 1998), and that bran alone accounts for 50% of the wheat milling by-products (Ash, 1992), this leaves about 65-85 million tons worldwide of wheat milling by-products, excluding bran. This figure is a rough estimate, since there are no consistent global statistics for individual categories such as wheat shorts, middlings, feed flour, red dog, and other remoulages, and that bran can be part of the final blends of wheat by-products.

Processes 

Wheat shorts, middlings and feed flour are already fine or floury fractions when they leave the mill, so they are commonly used as ingredients in mixed feeds without a separate processing step. The by-products can be mixed into mash feeds or pelleted as part of a compound feed; grinding and pelleting are optional feed-manufacturing treatments rather than steps required for every product (Blasi et al., 1998; Huang et al., 2014; Rosenfelder et al., 2013). Pelleting makes loose middlings denser, freer-flowing and less dusty, improving storage, transport and handling and reducing dust losses during loading and unloading (Blasi et al., 1998). Excessive grinding of already fine, low-fibre fractions such as feed flour or red dog can make the material sticky and may affect feed intake. During storage, the products should be kept dry and handled to avoid caking, bridging and spoilage, particularly during prolonged storage (Blasi et al., 1998; Huang et al., 2014).

Environmental impact 

The use of wheat milling by-products in animal feed may reduce the environmental impact of a ration when they replace other ingredients with higher impacts. However, they should not be considered impact-free: the wheat crop and its cultivation inputs are shared by flour and the milling co-products, and life-cycle assessments must assign these upstream burdens among the products (de Quelen et al., 2021; Keshavarz Afshar et al., 2026). The estimated footprint of a milling co-product depends substantially on how these shared burdens are allocated. In a recent US wheat-milling assessment, wheat flour received 87.7% of the upstream burden with economic allocation, compared with 73.0% with mass allocation and 72.2% with energy allocation; wheat bran and middlings received the remaining shares. Thus, a low footprint reported for a co-product may partly reflect the accounting method, as well as the product and process themselves (Keshavarz Afshar et al., 2026).

In a French pig-feed assessment, wheat middlings had a lower estimated climate-change impact per kilogram than wheat, and their inclusion in an environmentally optimized diet contributed to lower impacts for several categories. These results depended on the allocation method and the other ingredients in the diet; the locally sourced diet also increased land occupation. A consequential assessment found that shifting middlings from pig diets to dairy diets, while replacing them in pig diets with barley, reduced greenhouse-gas emissions by 329 kg CO2-eq per tonne of middlings when land-use-change emissions were included, but by only 9 kg CO2-eq per tonne when they were excluded. The environmental benefit therefore depends on what the middlings replace, their previous use, and the system boundary (de Quelen et al., 2021; van Zanten et al., 2014).

In a Brazilian wheat-flour life-cycle assessment, wheat cultivation was the main contributor to most assessed impacts, while milling made a comparatively small contribution overall, although it contributed more noticeably to freshwater eutrophication. This suggests that the crop-production system and allocation of its impacts are usually more important than the milling operation alone, but results will vary with location, energy sources, milling technology and product mix (Giongo et al., 2025).

Nutritional aspects
Nutritional attributes 

The chemical composition of wheat feed flour, shorts and middlings varies widely because each product contains different proportions of floury endosperm, bran and other mill streams. Across the Feedipedia table groups, dry matter averages about 88% as fed. Average crude protein ranges from 15.1% of dry matter in feed flour to about 17-18% in the middlings groups; ether extract averages about 2.7-4.6% and ash about 1.7-4.4% of dry matter. These are group averages, and individual samples vary around them (Feedipedia, 2026; Blasi et al., 1998; Slominski et al., 2004).

The ether extract represents a modest but variable lipid fraction, with the table-group means somewhat higher in middlings than in feed flour. In the limited fatty-acid data reported in the tables, linoleic acid is the largest fraction, followed by palmitic and oleic acids (Feedipedia, 2026).

Total mineral matter averages less in feed flour than in the middlings groups. Among the macro minerals reported, phosphorus and potassium are present at the highest concentrations; table-group mean values range from 4.4 to 9.0 g/kg dry matter for phosphorus and 6.2 to 11.3 g/kg for potassium. Mean calcium is lower (0.8-1.2 g/kg dry matter), while magnesium averages 1.8-3.3 g/kg. Sodium is present at much lower concentrations than these major minerals (Feedipedia, 2026).

The main compositional gradient is between starch and fibre. Feed flour is the most starch-rich and least fibrous group in the tables, with mean starch of 69.7% of dry matter and mean crude fibre of 1.1%. The middlings groups are classified by starch bands from less than 20% to more than 40%; their mean starch values range from 21.4% to 51.1%, while mean crude fibre ranges from 8.2% in the lowest-starch group to 6.5% in the highest-starch group. Mean neutral detergent fibre also declines across the middlings groups, from 36.2% to 29.4% of dry matter. Thus, the family spans floury, starch-rich material through substantially more fibrous products, although the bands overlap and are not sharply distinct feed types (Feedipedia, 2026).

This fibre–starch pattern reflects the proportions of endosperm and outer-kernel material retained in, or blended into, each product. Milling settings, flour extraction targets and the combination of mill streams can therefore produce substantial variation even when products share the same commercial name. In a Canadian analysis, wheat mill run samples likewise combined appreciable starch with high dietary fibre, illustrating that a named milling by-product can retain both floury and fibrous fractions (Blasi et al., 1998; Slominski et al., 2004; Rosenfelder et al., 2013).

These products can supply starch, protein, fat and fibre in a single feed ingredient, and their range of compositions can help match ingredients to different ration needs. Their advantages depend on identifying the material actually being purchased: variable composition and imprecise trade names make formulation from the product name alone unreliable, while fibre-rich batches may dilute dietary starch and energy concentration. Routine analysis and ration formulation around the actual batch are therefore important (Blasi et al., 1998; Rosenfelder et al., 2013; Woyengo et al., 2021).

Potential constraints 

Fibre and pentosans

The constraints associated with fibre depend on the milling fraction: in a comparison of wheat middlings and red dog fed to growing pigs, the more fibrous middlings had lower nutrient and energy digestibility than red dog (Casas et al., 2018). Arabinoxylans, also called pentosans, can restrict nutrient digestion in wheat millrun, middlings and shorts, and the extent to which xylanase alleviates this constraint differs among products (Nortey et al., 2008). Their effects are not explained solely by increased intestinal viscosity: in broiler experiments including wheat millrun, digesta viscosity was relatively low and did not consistently explain the responses to enzyme supplementation (Boros et al., 2004).

Phytate and sensitivity of phytase to heat

A substantial proportion of the phosphorus in wheat milling by-products is bound to phytate, so the total phosphorus concentration alone does not describe the amount available to pigs and poultry (Slominski et al., 2004; Casas et al., 2018). Wheat wheat milling by-products also contains endogenous phytase, which can release phosphorus from phytate. Steam pelleting can substantially reduce endogenous phytase activity, although some activity can survive; the activity measured in an unprocessed ingredient therefore cannot be assumed to remain unchanged in the pelleted feed (Slominski et al., 2004 ; Slominski et al., 2007).

Calcium and phosphorus imbalance

Wheat middlings generally contain much more phosphorus than calcium, and their contribution to the mineral balance must be considered when formulating diets containing substantial amounts of these products (Casas et al., 2018). In horses, phytic acid in pollard and bran can reduce calcium absorption and contribute to nutritional secondary hyperparathyroidism, or “big head disease”, when the ration does not provide an adequate calcium balance (Kohnke et al., 1999).

Moisture, moulds and storage

Pelleted wheat middlings are sensitive to moisture uptake: pellets can swell, soften and lose their ability to flow, causing storage and handling difficulties. Prolonged storage under warm, moist conditions can also favour the growth of storage moulds associated with heating and caking, even though pelleting initially reduces mould counts. Protection from water ingress and appropriate control of storage moisture are therefore important for preserving product quality (Blasi et al., 1998).

Mycotoxins

Wheat milling redistributes mycotoxins between fractions rather than destroying them. Contamination generally decreases in flour and semolina and increases in some feed by-products, but the distribution depends on the toxin, the location and severity of grain infection, and the milling process. The evidence is strongest for deoxynivalenol (DON), a Fusarium mycotoxin, although nivalenol, zearalenone, T-2 and HT-2 toxins are also relevant (Cheli et al., 2013). Shorts and middlings are not uniformly contaminated. In as study of industrial milling, DON concentrations in shorts or middlings were about 1.5–3.1 times those in the original grain, while flour shorts contained about 1.2–1.8 times the grain concentration. Other milling experiments found lower concentrations in fine middlings than in the grain. Thus, a single enrichment factor cannot be applied to this family of products (Cheli et al., 2013). In a more recent study of a commercial wheat batch, both shorts and red dog contained more DON than the whole kernels. Refined flour from the same batch also retained DON, although at a lower concentration, showing that floury fractions are not necessarily free of this contaminant (Guarino et al., 2025).

Sorting, cleaning and debranning can reduce contamination of the grain entering the mill by removing affected kernels and outer tissues, but can concentrate toxins in the rejected fractions. Protection of flour quality therefore does not establish the safety of the associated feed by-products; contamination should be assessed in the actual product intended for feeding (Cheli et al., 2013).

Ruminants 

Wheat middlings and related milling by-products are useful sources of energy and protein for ruminants. They can replace part of the cereal grain in balanced diets and provide supplements for forage-based feeding systems. Their feeding value depends on the proportions of flour, germ and fibrous tissues remaining after milling. Although middlings supply fermentable fibre, their small particle size limits its physical effectiveness, so their fibre cannot be considered equivalent to that of long forage (NASEM, 2021).

The protein and residual starch in middlings are readily available to rumen microorganisms: in an in situ comparison of eight feeds using three cannulated dairy cows, middlings had the highest estimated ruminal availability of both crude protein and starch. These measurements help explain the need to balance middlings with the other sources of fermentable energy and protein in the ration (Batajoo et al., 1998).

Inclusion rates

Published guidance and successful experimental inclusion levels are summarised below. Percentages refer to total diet dry matter (DM) unless another basis is specified. The sheep experiments tested particular durum-wheat middlings in balanced pelleted complete diets (Tufarelli et al., 2011a; Tufarelli et al., 2011b). Differences in starch and fibre content between milling products also limit the transfer of these results to other products, particularly starch-rich feed flour (NASEM, 2021).

 
Animal or feeding system Inclusion or allowance Basis and qualification
Lactating dairy cows 15–25% of total ration DM Published upper inclusion range, intended to prevent palatability problems; the guide also expresses this as 25–50% of the grain mix (Amaral-Phillips et al., 2006).
Lactating dairy cows: pelleted concentrate Up to 40% of the concentrate mixture Milk yield was maintained at this level, whereas 60% reduced yield despite unchanged intake. The concentrate made up about 60% of the ration; middlings percentages in its formulation were expressed as fed (Acedo et al., 1987).
Lactating dairy cows: partial forage replacement 15–17% of total diet DM Levels tested when replacing part of the maize silage or alfalfa contribution. Milk production was maintained or increased, but middlings fibre was less effective than forage fibre in stimulating rumination or supporting milk fat (Wagner et al., 1993; Depies et al., 1995).
Beef cattle No more than 50% of total diet DM Broad extension ceiling. Middlings should be combined with other ingredients because they ferment readily (Parish et al., 2021).
Growing and finishing beef cattle About 35.5% of total diet DM in growing diets; 35% and 50% in finishing diets Levels tested without significant changes in growth or feed efficiency; carcass traits were also unaffected in finishing steers. Ruminal pH was lower in a companion experiment with the 50% diet (ZoBell et al., 2003).
Lactating ewes 50% of total diet DM Successfully tested in a balanced pelleted total mixed ration; milk yield was maintained. (Tufarelli et al., 2011a).
Fattening lambs 60% of total diet DM Successfully tested for 50 days in a balanced pelleted total mixed ration (Tufarelli et al., 2011b).
Goats receiving forage Up to 1% of body weight daily as a supplement Published guidance for grazing goats; the guide does not specify the moisture basis (Gurung, 2013). A meat-goat trial tested 1% of body weight daily as fed, with hay available freely and the mineral balance corrected (Moore et al., 2002).

Dairy cattle

Middlings can help formulate dairy diets containing less cereal starch while supplying energy and rumen-degradable protein, but their fibre is less physically effective than forage fibre (NASEM, 2021). Their low calcium and high phosphorus contents require an appropriate mineral balance (Amaral-Phillips et al., 2006).

Two trials, each with 18 lactating cows, compared pelleted concentrates containing up to 40% or 60% middlings, replacing maize, sorghum grain and cottonseed meal. Concentrate supplied about 60% of the ration, with alfalfa hay providing the remainder. Milk yield was maintained with 20% or 40% middlings in the concentrate, but decreased at 60%; intake and milk fat percentage were unchanged. Thus, satisfactory intake of a high-middlings pellet did not necessarily indicate that it would maintain milk production (Acedo et al., 1987).

In a trial with 12 lactating Holstein cows, increasing middlings from 6.8 to 13.4% of diet DM, together with beet pulp and other formulation changes, reduced dietary starch from 24.6 to 17.7%. Feed intake, solids-corrected milk yield and feed efficiency were maintained, averaging 26.5 kg DM/day, 40.8 kg milk/day and 1.54 kg milk/kg DM, respectively. Ruminal pH and chewing activity were also unchanged, although organic matter digestibility was lower on the lower-starch diets (Dann et al., 2014).

Results were less favourable in a 10-week trial with 45 high-producing Holstein cows. Increasing middlings from 2.2 to 10.9% of diet DM, together with whole cottonseed replacing maize grain and soybean meal, reduced dietary starch from 27 to about 22% DM. Milk yield tended to decrease, milk protein content and yield declined, and feed conversion was poorer, despite greater DM intake relative to body weight in cows receiving the reduced-starch diet without amylase. These responses concern the combined ingredient substitution rather than an isolated effect of middlings (Ferraretto et al., 2011).

Middlings can also replace some forage. In a five-cow crossover experiment, diets containing 15% middlings DM and 38–39% maize silage were compared with a control containing 50% maize silage. Intake and fat-corrected milk production were higher on average with middlings, but ruminal pH tended to be lower. Calcium chloride added to the middlings diets lowered milk fat percentage, while sodium bicarbonate did not improve milk fat percentage, showing that the mineral formulation also affected the response (Wagner et al., 1993).

In a trial using 13 cows, a diet containing 17% middlings DM was compared with diets supplying additional fibre from alfalfa silage or ground maize cobs and with a low-fibre control. Intake and milk yield did not differ. Middlings increased milk fat percentage above the low-fibre control, but rumination was shorter than with additional alfalfa. Judged by the milk-fat response per unit of neutral detergent fibre, middlings fibre was approximately half as effective as alfalfa fibre (Depies et al., 1995).

Protein supplementation was examined in two trials with diets containing approximately 22–25% middlings DM. Cottonseed meal supported milk production similarly to soybean meal; substituting meat and bone meal produced variable responses in milk yield, with no improvement in energy-corrected milk. These historical comparisons illustrate the importance of the complete protein supplement, but did not test the effect of changing middlings inclusion itself (Bernard, 1997).

In a 10-week trial with 192 lactating cows, a ration containing 3.49% middlings DM, soybean meal and urea supported 43.6 kg/day of energy-corrected milk, similar to a ration containing rumen-protected soybean meal (43.7 kg/day). A third ration containing a fermentation by-product supported higher intake and milk production (45.3 kg/day). Because the protein supplements differed in several ingredients, this comparison demonstrates the use of middlings in a productive balanced ration rather than its individual effect on milk yield (Fessenden et al., 2020).

Beef cattle

Middlings provide both energy and protein for beef cattle, but their rapid fermentation and high phosphorus-to-calcium ratio require attention when formulating the ration (Parish et al., 2021). Replacing cereal grain with middlings does not necessarily increase ruminal pH (ZoBell et al., 2003).

In two 84-day growing trials, diets containing about 35.5% middlings DM replaced barley- or maize-based concentrates in maize silage and alfalfa-hay rations without significant changes in daily gain, intake or feed efficiency. In a subsequent 107-day trial with 24 finishing steers, diets containing 35% or 50% middlings DM maintained performance and carcass characteristics. However, in a companion digestion experiment, ruminal pH was lower with the 50% middlings diet than with the corn diet (5.55 versus 5.81), despite similar DM and acid detergent fibre digestibility (ZoBell et al., 2003).

For cattle consuming low-quality forage, the response to supplementation depends on the balance between fermentable carbohydrate and protein. In experiments with 28 yearling heifers and 60 winter-grazing beef cows, middlings supplied 1.7 kg DM/day as an intermediate-carbohydrate supplement, compared with soybean hulls plus soybean meal or barley plus soybean meal. Supplementation improved forage intake and digestion in the heifer experiment, but cow responses varied between years and increasing supplemental carbohydrate could depress forage utilisation. The benefits therefore depended on the basal forage and supplement balance (Bowman et al., 2004).

A more recent study evaluated middlings incorporated into wet maize distillers grains with solubles. In two short crossover experiments, each with four cattle, the blended product replaced unblended distillers grains at 40% of growing-diet DM or 30% of finishing-diet DM. Most measures of intake and digestibility were unchanged; the finishing diet containing the blend produced a higher ruminal pH and a small increase in DM digestibility. These percentages describe the whole blend, not middlings alone, and the study did not measure growth performance (Duncan et al., 2024).

Sheep

Durum-wheat middlings have successfully replaced substantial amounts of conventional concentrate ingredients in balanced pelleted diets for lactating ewes and fattening lambs. Their use at high inclusion rates is supported by specific feeding trials, rather than by a broadly established maximum allowance (Tufarelli et al., 2011a; Tufarelli et al., 2011b).

Forty lactating ewes received pelleted total mixed rations over 18 weeks, including two weeks of adaptation. A ration containing 50% middlings DM replaced a control containing 25.5% maize, with other ingredients adjusted to balance the diets. Milk yield, protein, lactose and clotting properties were maintained, while milk fat percentage and yield increased. A companion experiment with four adult rams found greater fibre digestibility but similar DM, organic matter and protein digestibility (Tufarelli et al., 2011a).

In a 50-day trial with 40 male Comisana lambs initially weighing about 13 kg, a pelleted ration reported to contain 60% middlings DM was compared with a control containing 40% maize. The diets were formulated to provide the same energy and protein concentrations. Daily gain increased from 174 to 199 g and feed conversion improved from 4.29 to 3.74 kg DM/kg gain. Cold carcass weight also increased, while most other carcass traits were unchanged. The middlings came from durum wheat of the Appulo cultivar (Tufarelli et al., 2011b).

Goats

Middlings are a useful energy and protein supplement for forage-fed goats, but their high phosphorus and low calcium contents limit unrestricted use and require mineral balancing (Gurung, 2013).

In a trial with 24 crossbred wether goats, initially weighing 27.4 kg, middlings, soybean hulls or maize gluten feed were supplied at 1% of body weight daily on an as-fed basis alongside freely available orchardgrass hay for 72 days. The control received hay supplemented with soybean meal, and the diets were adjusted to a calcium-to-phosphorus ratio of 1.5:1. Middlings maintained intake, daily gain and nutrient digestibility, while carcass weight was greater than in the control (15.6 versus 14.5 kg). Ruminal pH was lower with middlings (6.23 versus 6.52), but remained above 6 (Moore et al., 2002).

Pigs 

Wheat middlings, shorts and related milling by-products can replace part of the cereal grain and protein supplements in pig diets. Their usefulness depends largely on their fibre and starch contents: fibrous middlings dilute dietary energy, whereas more floury products have a higher energy value. Consequently, products sold under similar names cannot necessarily be used at the same inclusion rate (Casas et al., 2018; Stas et al., 2024).

Inclusion rates

The table distinguishes published feeding guidance from levels tested in particular trials. Percentages refer to the complete diet, as fed. These figures apply to the specified products and formulations. In particular, differences in fibre content between sources affect both energy value and amino acid digestibility (Espinosa et al., 2024).

 
Animal and product Inclusion Basis and qualification
Nursery pigs: middlings Up to 5% Conservative published guidance, rather than an experimentally established maximum (National Pork Board, 2008).
Nursery pigs: middlings 10–15% Range supported by the authors of four nursery trials. Responses varied between experiments, and formulation on a net energy basis was important (De Jong et al., 2014).
Piglets established on feed: millrun Up to 20% Successfully tested in balanced pelleted diets, starting two weeks after weaning. This result does not establish a maximum for diets fed immediately after weaning (Garcia et al., 2015b).
Growing and finishing pigs: middlings Up to 25% Published upper inclusion guidance (National Pork Board, 2008). Performance reductions have nevertheless occurred below this level in some formulations (Salyer et al., 2012).
Growing and finishing pigs: middlings 15% Tested in diets balanced for predicted net energy and digestible amino acids. Overall growth was maintained, but early growth and carcass yield were reduced; this is a tested level, not a maximum (Wu et al., 2016).
Gestating sows: middlings 30%, together with 15% soybean hulls Tested from days 35 to 104 of gestation, with feed allowances adjusted for dietary energy. A tested blend, rather than an established maximum for middlings alone (Lopez et al., 2021).
Lactating sows: middlings Up to 5% Conservative published guidance for inclusion in the complete diet (National Pork Board, 2008).

Growing pigs

Wheat milling by-products supply energy and amino acids, but increasing the proportion of fibrous milling fractions can reduce the amount of usable energy and digestible amino acids supplied by the feed. A comparison of ten middlings sources from Europe and the USA found substantial differences in these values, associated with differences in fibre content. Formulation therefore requires an appropriate estimate of the feeding value of the actual product (Espinosa et al., 2024). More floury products can have a higher feeding value than middlings. In a digestion trial with 12 growing pigs, red dog contained more starch and less fibre than ten sources of middlings, and supplied more digestible and metabolizable energy (Casas et al., 2018).

Piglets

In four experiments involving 780 nursery pigs, diets contained up to 20% middlings, with or without distillers grains. Increasing middlings reduced daily gain in two experiments, while another found unchanged daily gain but poorer feed efficiency at 20%. In the fourth experiment, soybean oil was added to some diets to equalise net energy. The authors concluded that 10–15% middlings could be used without impairing nursery performance, and that formulation on a net energy basis helped maintain performance relative to maize–soybean meal diets (De Jong et al., 2014).

Millrun was successfully used in 160 piglets initially weighing 9.8 kg. Pelleted wheat-based diets contained 0, 5, 10, 15 or 20% millrun, replacing part of the wheat and soybean meal, with canola oil and crystalline amino acids added to balance net energy and digestible amino acids. During the three-week trial, starting two weeks after weaning, daily gain and feed intake were unchanged. Daily gain averaged 509 and 510 g/day with 0 and 20% millrun, respectively, while feed efficiency increased from 0.61 to 0.65 kg gain/kg feed. The result concerns the complete reformulated diets, including their additional oil, rather than an isolated benefit of millrun (Garcia et al., 2015b).

Growing and finishing pigs

Middlings can reduce growth and carcass yield when their addition lowers dietary energy density. In a 62-day trial with 312 finishing pigs initially weighing about 64 kg, diets contained 0, 15 or 30% middlings without compensating for their lower energy value. Increasing inclusion reduced daily gain, feed efficiency, carcass weight and dressing percentage. Xylanase supplementation did not prevent these effects (Feoli et al., 2006).

Two further experiments, each with 288 growing-finishing pigs, examined middlings in diets also containing distillers grains. Increasing middlings to 20% reduced growth, feed efficiency, carcass weight and carcass yield. In the second experiment, added fat improved growth performance but did not eliminate the adverse effects of middlings on carcass yield or weight. Middlings also increased the iodine value of jowl fat in that experiment, indicating more unsaturated carcass fat (Salyer et al., 2012).

Balancing net energy gave more favourable growth results in a trial with 384 pigs, initially weighing 29 kg. Diets contained 0 or 15% middlings, with or without 30% distillers grains, and were balanced for predicted net energy and digestible amino acids. Middlings did not affect overall daily gain or feed efficiency, although daily gain was lower during the first phase, from about 29 to 50 kg. Carcass yield decreased from 73.1 to 72.7%, showing that maintained live-weight performance did not fully preserve carcass output (Wu et al., 2016).

Sows

The fibre and bulk of middlings can be useful in gestation diets, but feed allowances must supply sufficient energy and nutrients. Where high inclusion dilutes dietary energy, the daily allowance needs to be increased accordingly. More restrictive inclusion guidance is given for lactation, when nutrient intake must support milk production (National Pork Board, 2008).

Gestation

A trial with 180 group-housed sows compared a diet containing 30% middlings and 15% soybean hulls with one containing 30% distillers grains and 30% maize germ meal. The diets were fed from days 35 to 104 of gestation, with different daily allowances to account for their energy contents; all sows subsequently received the same lactation diet. Sows receiving the middlings–soybean hull diet had less backfat and weaned lighter piglets than those receiving the other fibrous diet. Aggression was influenced mainly by the length of the feeding partitions (Lopez et al., 2021).

Farrowing and lactation

A conference abstract reported a trial with 860 sows receiving a maize top-dressing or a 50:50 middlings–soybean hull blend, with or without a feed additive intended to stimulate fibre fermentation. The top-dressing supplied 454 g/day (227 g middlings/day in the blend), starting one day after entry into farrowing accommodation until weaning. The blend reduced pre-farrowing constipation by 27% relative to the control and reduced the frequency of absent or dry, pellet-like faeces after farrowing by 22%. Colostrum Brix readings were unchanged, and the additive did not improve constipation outcomes (Self et al., 2025).

Poultry 

Wheat middlings, shorts and related milling by-products can supply energy and protein in poultry diets, but their value depends on the proportions of floury and fibrous milling fractions. Compared with wheat grain, millrun contains more non-starch polysaccharides and less starch, so its inclusion requires an appropriate allowance for its lower energy value (Slominski et al., 2004). Formulation on a digestible amino acid basis also helps account for differences in the availability of protein from feed ingredients (Adedokun et al., 2015).

Bird type and the method used to measure digestibility affect the interpretation of feeding values. One study found higher apparent ileal amino acid digestibility of middlings in laying hens than in broilers (Huang et al., 2007). However, another found no difference between broilers and hens in standardized ileal amino acid digestibility of the middlings sample tested (Adedokun et al., 2015).

Inclusion rates

The following figures describe tested inclusions and experimental feed formulations. They are not universal maximum recommendations. Percentages refer to the complete diet, as fed. Results for a particular millrun, middlings or pollard should not be transferred directly to another milling product; in addition to ingredient composition, diet formulation and processing influence the response (Slominski et al., 2004; Boros et al., 2004).

 
Bird and product Inclusion Basis and qualification
Broilers: middlings 5% Tested as a pellet binder from 11 to 49 days; growth and feed conversion improved. Not a maximum inclusion trial (Saleh et al., 2021).
Broilers: millrun 10% Maintained growth and feed conversion in a short mash-feed trial from 4 to 18 days (Boros et al., 2004).
Broilers: middlings with sunflower meal 10% in starter feed; 18.5% in finisher feed Tested together with 7.5% and 8% sunflower meal, respectively. Untreated ingredients maintained growth, but replacing most of them with enzyme-pretreated material reduced growth. The trial does not isolate middlings effects (Muchiri et al., 2022).
Laying hens: middlings 20–30% Tested from 44 to 51 weeks of age, with dietary energy adjustments and enzyme treatments. Laying rate and egg mass were maintained, but feed conversion was poorer than in the control. These are tested levels, not performance-neutral recommendations (Abudabos, 2011b).
Turkey poults: middlings 10% Tested in balanced wheat-based crumbles to 28 days; growth was maintained. No upper limit was established (Njeri et al., 2023).
Breeding Japanese quails: middlings 4.3% Basal-diet inclusion in a hatchability experiment; middlings level was not a treatment (Ayaşan, 2013).
Pekin ducks: middlings 8% in starter feed; 14% in grower feed Basal-diet levels in an enzyme trial, not tested maxima (Hong et al., 2002).

Broilers

Moderate amounts of millrun can replace part of the wheat in broiler diets without impairing growth, although the evidence concerns particular products and formulations (Boros et al., 2004). Middlings can also contribute to the physical quality of pelleted feeds (Saleh et al., 2021).

In a two-week trial, broilers received mash diets containing 10% millrun in place of wheat from 4 to 18 days of age. Weight gain and feed conversion were similar to those of birds receiving the wheat–soybean meal control. The clearest growth response to enzymes in the same paper concerned wheat screenings, while its longer production trial used screenings, barley and canola meal; those responses should not be attributed specifically to millrun (Boros et al., 2004).

In a trial with Egyptian broilers, 5% middlings in grower and finisher diets from 11 to 49 days increased the proportion of intact pellets and reduced fines. Weight gain increased and feed conversion improved compared with the control. The result supports a useful role at a low inclusion, including an effect on feed structure, rather than establishing a maximum replacement of cereal grain (Saleh et al., 2021).

Enzyme treatment does not always improve performance. In a 42-day trial with 288 broilers, diets contained 10% middlings and 7.5% sunflower meal in the starter phase, and 18.5% middlings and 8% sunflower meal in the finisher phase. Increasing proportions of both ingredients were replaced by material pretreated with fibre-degrading enzymes, incubated and oven-dried. Untreated ingredients and replacement of up to half with pretreated material maintained weight gain relative to the maize–soybean meal control, whereas 75% or complete replacement reduced growth. These were lower-energy diets, and the response cannot be separated into effects of middlings, sunflower meal and their processing (Muchiri et al., 2022).

Laying hens

Middlings can replace substantial amounts of cereal grain in layer diets while maintaining the laying rate, but feed intake, feed conversion and egg size may change. Consequently, egg production alone is insufficient to judge their feeding value (Bai et al., 1992; Abudabos, 2011b).

Long-term experiments compared a maize–soybean meal diet with a diet containing 25% middlings in several commercial strains. Laying rate was maintained, while egg weight, feed intake and feed required for egg production increased. In a separate 44-week comparison, hens receiving 89% middlings laid fewer eggs than those receiving 45%: hen-day production averaged 67.5% versus 79%. Feed and water intakes were also higher at 89%. These extreme inclusions illustrate the consequences of relying heavily on middlings, rather than practical feeding recommendations (Bai et al., 1992).

In a seven-week trial with 140 hens from 44 to 51 weeks of age, diets containing 20 or 30% middlings were compared with a maize–soybean meal control. The middlings diets were formulated at the control's calculated energy level, with or without a combination of xylanase, amylase and protease, or at a slightly lower energy level with enzymes. Feed intake, laying rate and egg mass were unaffected, but feed conversion was poorer: averages were 1.87 and 1.95 kg feed/kg egg mass at 20 and 30% middlings, respectively, compared with 1.80 for the control. Enzymes increased egg weight in the normal-energy diet containing 20% middlings, but did not significantly improve feed conversion or measured metabolizable energy. Thus, maintained egg output did not imply equal feed efficiency (Abudabos, 2011b).

A more recent organic-farm trial with 80 hens used 36.3% of a mixture of spelt and emmer middlings in balanced diets. Laying rate and egg mass were maintained, and feed conversion improved, but eggs were smaller and their shells were thinner and weaker. These findings concern the particular ancient-wheat milling products tested (Lombardi et al., 2020).

Turkeys

Turkey poults have tolerated middlings in balanced diets, but the trial below does not establish an upper inclusion rate for growing or breeding turkeys (Njeri et al., 2023).

An experiment with 480 turkey poults compared balanced wheat diets containing 0 or 10% middlings, alongside maize diets and a multienzyme treatment. Through 28 days, added fibre did not affect weight gain or feed conversion; middlings increased gizzard weight in wheat-fed poults. Enzymes increased metabolizable energy in wheat diets without middlings, but not those containing them (Njeri et al., 2023).

Quails

Published use of middlings is documented in breeding-quail feed, but the evidence reviewed here does not establish a quail-specific optimum or maximum inclusion.

A six-week hatchability experiment with Japanese quails used a basal diet containing 43.43 g middlings/kg feed (4.34%). Both treatment groups received this inclusion; the experimental comparison concerned a plant extract. Thus, the study documents the use of middlings in a complete breeder diet, but does not measure their effect on laying performance, fertility or hatchability (Ayaşan, 2013).

Ducks

Ducks can use middlings-containing diets, but the available enzyme trial does not establish the ingredient’s independent effect. In a trial with 360 male Pekin ducks, mash diets based on maize and soybean meal contained 7.93% middlings in the starter phase and 13.95% in the grower phase. A combination of amylase, protease and xylanase increased weight gain by 6–8% and improved feed conversion over the six-week feeding period. Amino acid utilization improved, but measured metabolizable energy was unchanged. As the experiment compared enzyme additions, it did not establish a maximum middlings inclusion (Hong et al., 2002).

Rabbits 

Wheat middlings, shorts and related milling by-products can supply energy and protein in complete rabbit feeds. Their usefulness depends on the balance between starch and fibre: flour-rich products contribute more energy, while more fibrous products can replace some cereal grain. They should be combined with ingredients that provide sufficient structural fibre and a balanced supply of amino acids and minerals (de Blas et al., 2020).

Inclusion rates

The levels below distinguish experimental formulations from published formulation guidance. Percentages refer to the complete formulated feed, except where a dry matter (DM) basis is specified. Trials combining middlings and bran do not establish the maximum inclusion of middlings alone, and their results should not be transferred directly to starch-rich feed flour.

Animal or feeding system Inclusion Basis and qualification
Growing rabbits: durum-wheat middlings 23%, together with 30% wheat bran Tested from 28 to 84 days. Feed conversion improved, but growth was slower than in the control; the authors requested further work on digestive safety. An experimental level, not a recommended maximum (Berchiche et al., 2000).
Growing rabbits: durum-wheat middlings 10%, together with 50–57% wheat bran Tested from 28 to 77 days. Growth and feed conversion were maintained, but substantial mortality occurred across all diets. The 60–67% figure for total wheat by-products must not be interpreted as middlings inclusion (Lakabi-Ioualitene et al., 2008).
Adult does at maintenance and during lactation 9.35% of total diet DM Calculated from 18.7% middlings in a concentrate supplying half the diet DM; chopped grass hay supplied the remainder. A basal formulation in a four-doe physiology experiment, not an inclusion-response trial (Findeisen et al., 2021).
Compound feeds for pet rabbits: wheat feed Suggested maximum of 30% The guide also restricts all wheat products combined to 30% of the compound feed. This is guidance for pet-rabbit compounds, not an established ceiling for meat rabbits or 30% of a hay-plus-concentrate ration (Lowe, 2020).

Growing rabbits

Middlings can contribute to diets based largely on locally available milling by-products, but satisfactory feed conversion does not by itself demonstrate an adequate fibre balance or digestive safety (Berchiche et al., 2000; Lakabi-Ioualitene et al., 2008).

In an eight-week trial with 81 rabbits of a local Algerian population, a pelleted diet containing 23% durum-wheat middlings and 30% bran replaced all the barley and part of the soybean meal in the control. Feed conversion improved from 4.52 to 3.91, but daily gain decreased from 30.5 to 28.8 g and final live weight was lower. Hot carcass weight was unchanged. Four of the 27 rabbits receiving this diet died with diarrhoea; the authors recommended increasing indigestible fibre and conducting further trials before wider use (Berchiche et al., 2000).

A subsequent trial with 81 rabbits compared a conventional diet with two diets containing 10% durum-wheat middlings and either 50% or 57% bran, replacing barley and most or all of the soybean meal. Between 28 and 77 days, daily gain, feed conversion and cold-carcass dressing percentage did not differ significantly, despite lower dietary energy digestibility. However, mortality was high across treatments, at about 23%. The authors considered insufficient fibre a possible contributing factor and called for trials with diets meeting current fibre recommendations (Lakabi-Ioualitene et al., 2008).

Breeding does

Middlings have been used in feeds for lactating does, but the available experiment does not determine their optimum inclusion or their effect on reproductive performance (Findeisen et al., 2021).

Four adult does received a constant mixture of equal DM proportions of chopped grass hay and concentrate, the latter containing 18.7% middlings. Thus, middlings supplied 9.35% of total diet DM. The ration was completely consumed at both maintenance and lactation feeding levels. Higher intake during lactation shortened digesta retention and reduced DM and fibre digestibility, while protein digestibility and chewing efficiency were maintained. These results describe the response to intake of the whole ration, rather than a specific effect of middlings (Findeisen et al., 2021).

Pet rabbits

Low-starch wheat feed is a useful ingredient in prepared pet-rabbit feeds, provided that the complete formulation supplies adequate fibre. The suggested 30% limit applies to the compound feed and includes the contribution of other wheat products. Hay or fresh long fibre should still be freely available, and the amount of prepared feed should be adjusted to body condition (Lowe, 2020).

Horses and donkeys 

Wheat middlings, shorts and feed flour can provide energy and protein in balanced concentrates for equids. Their feeding value varies with the proportions of flour and fibrous milling fractions. Their phosphorus-rich, calcium-poor mineral balance and, particularly in flour-rich products, their starch contribution limit unrestricted use (Martin-Rosset, 2015).

Inclusion rates

Published allowances concern different products and feeding situations, so they should not be treated as interchangeable limits. In particular, an allowance for wheat milling by-products collectively does not establish an optimum for an individual middlings or feed-flour sample.

 
Animal or product Guidance Basis and qualification
Horses: wheat milling by-products collectively Do not exceed 20–30% of the diet General guidance intended to avoid worsening the calcium-to-phosphorus balance, particularly in grain-rich diets. The passage does not specify the moisture basis or establish separate limits for middlings, shorts and feed flour (Martin-Rosset, 2015).
Horses: pollard used routinely as a conditioning supplement 5% of the total ration, or approximately 100 g/100 kg body weight daily Older Australian practical guidance for routine use of pollard. It addresses a different feeding situation from the broader INRA allowance and is not an experimentally established maximum (Kohnke et al., 1999).
Donkeys No product-specific maximum established The sources reviewed do not provide a validated inclusion limit for clearly identified wheat middlings, shorts or feed flour in donkey diets.

Horses

Middlings can supply protein and energy in compound feeds, while feed flour provides more starch and generally more energy. Formulation should account for these differences and correct the mineral balance; the milling by-products should not be assigned a common feeding value merely because they share a commercial name (Martin-Rosset, 2015).

Australian guidance describes wheat pollard as a conditioning feed, but notes that it is dusty and should be dampened and mixed into the feed. Its energy contribution can promote weight gain, so the allowance should be matched to the horse's requirements and body condition (Kohnke et al., 1999).

In a study allocating 30 adult horses to forage alone or forage supplemented with maize, oats or pelleted wheat middlings, the supplements were introduced gradually over two weeks. Middlings containing 23.9% starch on a DM basis increased starch-utilizing bacteria and reduced cellulose-degrading bacteria in faeces. Faecal pH decreased from about 7.8 to 6.5, although no faecal lactate was detected. This demonstrates that a relatively fibrous middlings product can still alter hindgut fermentation. The experiment measured microbial and faecal responses, rather than growth or exercise performance, and did not establish a maximum middlings inclusion (Harlow et al., 2016).

Donkeys

The potential role of middlings in donkey feeding is as an ingredient of a balanced supplement when forage alone cannot meet requirements. Donkeys have lower maintenance energy expenditure and generally use poor-quality forage more efficiently than horses, so horse concentrate allowances should not be transferred directly. Supplementation should reflect work, lactation, forage quality and body condition while maintaining the forage contribution (Martin-Rosset, 2015).

No feeding trial that clearly identified middlings, shorts or feed flour and tested their inclusion response in donkeys was found in the sources reviewed. Consequently, a donkey-specific benefit or maximum allowance cannot presently be stated.

Fish 

Wheat middlings, shorts and related milling by-products can provide energy and some protein in fish feeds, particularly for omnivorous freshwater species. Their value depends on the milling fraction, its fibre and starch contents, the processing of the feed, and the species being fed. They should be incorporated into balanced diets rather than treated as direct replacements for protein concentrates.

A relatively good protein digestibility does not necessarily indicate a high energy value: in Nile tilapia, the protein in middlings was well digested, while dry matter and energy digestibility were substantially lower than those of wheat grain. Fibre and ash contents were negatively associated with nutrient and energy digestibility across the wheat products tested (Vidal et al., 2017).

Inclusion rates

The following figures distinguish published formulation guidance from experimental inclusions. They are percentages of the complete feed; a dry matter (DM) basis is specified where reported by the source. Successful use of one product does not establish a maximum for other milling fractions or fish species.

Published guidance and experimental inclusions of wheat middlings in fish feeds
Fish or feeding system Inclusion Interpretation
Fish feeds: general guidance 10–20% Older handbook guidance for pollard or middlings, without a species-specific maximum or a stated moisture basis (Hertrampf et al., 2000).
Channel catfish Up to about 25%; sometimes 10–15% under very humid conditions Practical guidance for feed manufacture and feeding. Higher levels can cause sticky feed and pellet clumping; this is not a toxicity threshold (Robinson et al., 2001).
Channel catfish: pond trials 6.15–16.24% as fed in 32% protein diets; 20% as fed in 28% protein diets Tested in extruded floating feeds over two growing seasons in each of two experiments. Middlings had no significant effect on growth or feed conversion; the study did not establish a maximum (Li et al., 2010).
Common carp juveniles 41.1–41.7% of diet DM Present in all diets of a 56-day protein-source trial. Demonstrates use in a balanced formulation, but does not establish an optimum or maximum for middlings (Fan et al., 2024).
Nile tilapia 30% in digestibility test diets; 7% of diet DM in a growth trial The digestibility test was not a growth trial. At 7%, growth was lower after part of the fish meal was replaced, but the test diet also contained less protein; this does not establish a 7% tolerance limit (Vidal et al., 2017; Niyibizi et al., 2023).
Black pacu / tambaqui 30% of diet DM Tested for 24 weeks, with growth, feed conversion and survival similar to diets containing alternative carbohydrate sources (Lochmann et al., 2009).
Turbot 24.76% of diet DM Replaced the wheat flour in a fish-meal-based diet for nine weeks without significant effects on growth or feed conversion. A tested inclusion, not an established maximum (Sevgili et al., 2021).
Juvenile red drum 28.0–63.3% of diet DM Experimental range in a lipid–carbohydrate study. The diet with 28% gave the best growth; higher carbohydrate levels reduced performance. These levels are not recommendations (Ellis et al., 1991).

Salmonids (Oncorhynchus kisutch and Oncorhynchus mykiss)

For salmonids, middlings can supply digestible protein, but their mineral content should not be equated with an equivalent supply of available minerals. In experiments in the USA with coho salmon (Oncorhynchus kisutch) and rainbow trout (Oncorhynchus mykiss), apparent protein digestibility exceeded 80% for all ingredients tested, including wheat middlings and flour, whereas mineral availability varied considerably. The diets contained 30% test ingredient, but the experiment measured digestibility and mineral absorption rather than long-term growth, and therefore does not establish a recommended inclusion ceiling (Sugiura et al., 1998).

Common carp (Cyprinus carpio)

Wheat middlings can form a substantial part of balanced common carp (Cyprinus carpio) feeds, although evidence from formulations containing a fixed amount of middlings does not determine their individual feeding value or maximum inclusion. In a 56-day trial in China, common carp initially weighing 4.17 g received pelleted diets containing 41.1–41.7% middlings on a DM basis, together with soybean meal, soybean protein concentrate, rapeseed meal and different proportions of fish meal and cottonseed protein concentrate. The diets contained about 30.5% crude protein. Growth and feed utilization were best when cottonseed protein concentrate replaced half the fish meal. Since middlings were included at almost the same level in every diet, the differences reflect the protein-source comparison and cannot be attributed to middlings (Fan et al., 2024).

Tilapia (Oreochromis niloticus)

Middlings can contribute protein and carbohydrate to tilapia feeds, but their fibre content can restrict the amount of digestible energy supplied. In an experiment in Brazil with extruded test diets for Nile tilapia (Oreochromis niloticus), middlings containing 8.5% crude fibre on a DM basis had apparent digestibility coefficients of 84.2% for protein, 57.5% for energy and 55.9% for DM. Their energy digestibility was lower than that of either wheat grain or the bran sample tested, illustrating the need to characterize the actual product rather than infer feeding value from its name (Vidal et al., 2017).

In a 70-day trial in Rwanda, a diet containing 7% middlings on a DM basis replaced 7 percentage points of fish meal in a mixed-ingredient control diet. Final weight was lower, at 50.0 versus 60.2 g, while survival did not differ significantly. However, the analyzed protein content was also lower in the middlings diet, at 24.1 versus 29.8% of DM. Thus, the comparison shows that this replacement strategy was nutritionally inadequate, rather than demonstrating that tilapia cannot tolerate 7% middlings (Niyibizi et al., 2023).

Channel catfish (Ictalurus punctatus)

Wheat middlings are a practical alternative to part of the maize in channel catfish (Ictalurus punctatus) feeds and can improve pellet binding. US guidance describes routine use up to about 25%, with lower levels sometimes necessary under very humid manufacturing conditions because of pellet stickiness and clumping (Robinson et al., 2001).

Two pond experiments in the USA compared extruded floating feeds containing different combinations of soybean meal, maize, cottonseed meal, middlings and fish meal. In the first experiment, middlings were included at 6.15–16.24% as fed in diets formulated to contain 32% protein; in the second, they were included at 20% as fed in diets formulated to contain 28% protein. Fish initially weighing 34 and 49 g, respectively, were stocked at 18,530/ha and fed over two growing seasons in each experiment. Comparisons of diets with and without middlings showed no significant effect on weight gain or feed conversion. Survival, processing yields and fillet composition did not differ significantly among dietary treatments. These results support the tested inclusions in balanced pond feeds, but do not establish an optimum or maximum (Li et al., 2010).

Silver catfish (Rhamdia quelen)

A 45-day trial in Brazil with 288 juvenile silver catfish (Rhamdia quelen), initially weighing about 12.8 g, tested wheat middlings (in the original Portuguese, the product is named triguilho, a term also used for off-grade wheat containing fragmented or poorly filled grains, so there may be a possible confusion). It replaced 0, 33, 67 or 100% of the maize, corresponding to actual inclusions of 0, 14.42, 28.84 and 43.26% in the diet. Diets were extruded, balanced for calculated digestible protein and energy, and supplemented or not with 1,500 FTU phytase/kg. Growth, feed conversion and survival did not differ significantly, while the highest replacement reduced visceral fat deposition. Phytase did not significantly improve growth performance (Pessini et al., 2020).

Black pacu (Colossoma macropomum)

Middlings can be used as a carbohydrate source for black pacu or tambaqui (Colossoma macropomum), although their higher fibre content may reduce their value relative to starch-rich local feedstuffs. In a 24-week trial at Southern Illinois University in the USA, a diet containing 30% middlings on a DM basis was compared with diets containing 30% cassava root, peach-palm fruit or plantain fruit. Final weight, weight gain, feed conversion and survival did not differ significantly. Final weight was about 539 g with middlings and 527–559 g with the alternatives. Protein efficiency was lower with middlings, but that diet also contained more protein, at 31.8 versus 27.0–28.1% of DM (Lochmann et al., 2009).

Turbot (Scophthalmus maximus)

Middlings can replace wheat flour in a protein-rich turbot (Scophthalmus maximus) diet, but this result concerns one milling product and a relatively large juvenile fish. In a nine-week trial in Turkey with fish initially weighing about 267 g, 24.76% middlings on a DM basis replaced all the wheat flour in a fish-meal-based control diet. Diets contained about 54% protein and 13–14% lipid on a DM basis. Final weight, feed intake, feed conversion, protein efficiency and nutrient retention did not differ significantly from the control. Final weights were about 406 and 413 g, and feed conversion ratios were 0.95 and 0.87, respectively. This supports the tested replacement, but does not establish an upper inclusion limit or demonstrate a benefit from additional phosphorus (Sevgili et al., 2021).

Red drum (Sciaenops ocellatus)

High middlings inclusions require caution in juvenile red drum (Sciaenops ocellatus), whose ability to use carbohydrate as an energy source is more limited than that of warm-water omnivorous fish. In an eight-week trial in the USA, fish initially weighing 2.9 g received six diets containing about 32% protein, two lipid levels and increasing carbohydrate levels, with 28.0–63.3% middlings on a DM basis. The best weight gain was obtained with 28% middlings, about 10% lipid and 24% nitrogen-free extract. Growth, feed efficiency and protein and energy retention declined as dietary carbohydrate increased, while survival did not differ significantly. Middlings, fish meal, oil and cellulose filler all varied between diets, so the experiment does not isolate an ingredient-specific maximum (Ellis et al., 1991).

Crustaceans 

Wheat milling by-products can provide carbohydrate and some protein in crustacean feeds. The distinction between fibrous middlings and low-fibre feed flour is particularly important: clear flours can supply highly digestible energy, while their usefulness as feed ingredients also depends on the physical quality of the pellets (Nieto-López et al., 2011).

Water stability should be considered together with ingestion. In a trial using middlings-containing shrimp feeds, a binder that reduced dry matter loss also produced harder wet pellets and lower feed consumption. Thus, greater pellet stability alone did not indicate a better feeding response (Cerecer-Cota et al., 2005).

Inclusion rates

The available evidence does not establish a single maximum for prawns or shrimps. Older general guidance and experimental formulations are distinguished below. Flour inclusions should not be transferred directly to more fibrous middlings.

 
Animal and product Inclusion Interpretation
Crustacean feeds: middlings or pollard 3–5% Older general handbook guidance, without a species-specific maximum or stated moisture basis. The same handbook gives different allowances for wheat flour (Hertrampf et al., 2000).
Amazon River prawn: middlings 24.4–24.5% of diet DM Present in every diet of a 60-day yeast-additive experiment; not an inclusion-response trial (Hisano et al., 2008).
Pacific white shrimp: middlings 16.5–20% as mixed Present in commercial-type steam-pelleted diets in a 28-day binder experiment; not a maximum established by a growth trial (Cerecer-Cota et al., 2005).
Pacific white shrimp: clear flours 30% test ingredient Digestibility measurements with hard-wheat clear flour and mixed-wheat second clear flour, not a growth-based recommendation (Nieto-López et al., 2011).
Giant tiger shrimp: clear flours 35% of the formulated feed First- and second-grade clear flours tested for eight weeks with growth and feed conversion comparable to straight flour. This applies to the low-fibre flours tested (Shiau et al., 1991).

Amazon River prawn (Macrobrachium amazonicum)

Middlings have been used at substantial levels in complete feeds for Amazon River prawn (Macrobrachium amazonicum), but the available trial does not determine their optimum inclusion or compare them with another carbohydrate source. In a 60-day trial in Brazil, juvenile prawns initially weighing 1.27 g received pelleted diets containing 24.4–24.5% middlings on a DM basis, together with about 32% soybean meal and 28% fish meal. Diets contained about 35% protein and were supplemented or not with yeast or yeast derivatives. Survival was 92.5–97.5% and did not differ significantly between treatments. Mannan-oligosaccharide supplementation improved weight gain, while feed conversion did not differ significantly. These results document the use of middlings in the basal formulation; the growth response was to the additive and cannot be attributed to middlings (Hisano et al., 2008).

Pacific white shrimp (Litopenaeus vannamei)

For Pacific white shrimp (Litopenaeus vannamei), low-grade milling flours can provide digestible carbohydrate, while middlings must also be assessed for their effects on pellet properties and ingestion (Nieto-López et al., 2011; Cerecer-Cota et al., 2005).

In a digestibility experiment in Mexico, hard red winter clear flour and a second clear flour obtained from milling medium-hard and soft wheats were included at 30% in test diets. Apparent energy digestibility was about 96% for both clear flours, compared with 83–86% for the whole-grain meals and semolina tested. Protein digestibility did not differ significantly between products. These results support the energy value of the flour-rich fractions, but the experiment did not establish their effect on long-term growth or a maximum inclusion rate (Nieto-López et al., 2011). A separate 28-day experiment used commercial-type steam-pelleted diets containing 16.5–20% middlings, 30% whole-wheat meal and 20% fish meal. Different binders replaced small amounts of middlings. A synthetic binder reduced DM loss after one hour in seawater from 10.4 to 2.6%, but increased wet-pellet hardness and reduced feed consumption. The study demonstrates the importance of checking both water stability and consumption in middlings-containing formulations; it does not establish an independent growth benefit or inclusion ceiling for middlings (Cerecer-Cota et al., 2005).

Giant tiger shrimp (Penaeus monodon)

First- and second-grade clear flours can be suitable alternatives to straight wheat flour for giant tiger shrimp (Penaeus monodon), provided that the material is a comparable low-fibre flour. In an eight-week trial in Taiwan, juveniles initially weighing about 1.03 g received diets containing 35% straight flour, first-grade clear flour or second-grade clear flour. The clear flours contained less than 1% crude fibre. Growth, feed conversion and apparent nutrient digestibility did not differ significantly among these three flour treatments. The flour diets also had similar water stability, with about 12–13% DM loss after two hours of immersion. This finding concerns milling flours and should not be interpreted as support for 35% fibrous middlings in shrimp feeds (Shiau et al., 1991).

Nutritional tables

Avg: average or predicted value; SD: standard deviation; Min: minimum value; Max: maximum value; Nb: number of values (samples) used

Main analysis Unit Avg SD Min Max Nb  
Dry matter % as fed 87.6 1.4 83.8 96.3 506  
Crude protein % DM 15.1 2.2 8.4 20.5 514  
Crude fibre % DM 1.1 0.6 0.1 3.3 107 *
Neutral detergent fibre % DM 8.2 3.6 5 15.8 8 *
Acid detergent fibre % DM 1.7 1 0.4 3.2 7 *
Lignin % DM 0.5 0.1 0.3 0.8 10  
Ether extract % DM 2.7 0.9 0.7 4.4 107 *
Ash % DM 1.7 0.6 0.5 3.1 152 *
Insoluble ash % DM 0.07       1  
Starch (polarimetry) % DM 69.7 7.3 51.1 83.2 463  
Starch (enzymatic) % DM 62.5         *
Total sugars % DM 5.6         *
Gross energy MJ/kg DM 18.9 1 16.2 20.8 16 *
               
Amino acids Unit Avg SD Min Max Nb  
Alanine g/16g N 4.2         *
Arginine g/16g N 5.7 0.7 4 5.7 5 *
Aspartic acid g/16g N 6         *
Cystine g/16g N 2 0.1 2 2.3 6 *
Glutamic acid g/16g N 23.7         *
Glycine g/16g N 4.8         *
Histidine g/16g N 2.7   2.2 2.7 4 *
Isoleucine g/16g N 3.2 0.3 3.1 3.8 5 *
Leucine g/16g N 6.3 0.3 5.9 6.6 5 *
Lysine g/16g N 3.7 0.3 2.8 3.9 7 *
Methionine g/16g N 1.5 0.2 1.4 1.9 7 *
Methionine+cystine g/16g N 3.5         *
Phenylalanine g/16g N 4.1 0.2 4 4.4 5 *
Phenylalanine+tyrosine g/16g N 6.9       1 *
Proline g/16g N 6.7       1 *
Serine g/16g N 4.3 0.2 4.1 4.7 5 *
Threonine g/16g N 3.1 0.1 2.8 3.1 7 *
Tryptophan g/16g N 1.2         *
Tyrosine g/16g N 2.8       1 *
Valine g/16g N 4.5 0.4 3.8 4.7 5 *
               
Fatty acids Unit Avg SD Min Max Nb  
Myristic acid C14:0 % fatty acids 0.1   0 0.3 4  
Palmitic acid C16:0 % fatty acids 17.8 0.9 15.5 18.3 9  
Palmitoleic acid C16:1 % fatty acids 0.4          
Stearic acid C18:0 % fatty acids 0.8 0.07 0.7 0.9 8  
Oleic acid C18:1 % fatty acids 15.2 2.7 11.3 20.4 11  
Linoleic acid C18:2 % fatty acids 56.4          
Linolenic acid C18:3 % fatty acids 5.9 1 5.4 8.5 9  
               
Minerals Unit Avg SD Min Max Nb  
Calcium g/kg DM 0.8 0.4 0.8 2.1 8 *
Phosphorus g/kg DM 4.4 1.3 1.8 5.8 12 *
Potassium g/kg DM 6.2       1 *
Sodium g/kg DM 0.07   0 0.17 4  
Chlorine g/kg DM 0.7       1  
Magnesium g/kg DM 1.8          
Sulfur g/kg DM 1.7       1  
Manganese mg/kg DM 57          
Zinc mg/kg DM 45          
Copper mg/kg DM 6          
Iron mg/kg DM 16   15 16 2  
Selenium mg/kg DM 0.2          
               
Pig nutritive values Unit Avg SD Min Max Nb  
Energy digestibility, growing pig % 92.7         *
DE growing pig MJ/kg DM 17.5         *
MEn growing pig MJ/kg DM 17         *
NE growing pig MJ/kg DM 13.1         *
Nitrogen digestibility, growing pig % 87.1         *
               
Poultry nutritive values Unit Avg SD Min Max Nb  
AMEn cockerel MJ/kg DM 14.7         *
AMEn broiler MJ/kg DM 14.1       1 *
               
Ruminants nutritive values Unit Avg SD Min Max Nb  
OM digestibility, ruminants % 86.8         *
Energy digestibility, ruminants % 84.6         *
ME ruminants MJ/kg DM 13.3         *
Nitrogen digestibility, ruminants % 71         *
Nitrogen degradability (effective, k=6%) % 76         *
Nitrogen degradability (effective, k=4%) % 80         *
a (N) % 35   32 56 2  
b (N) % 57   39 83 3  
c (N) h-1 0.155   0.134 0.182 3  
Dry matter degradability (effective, k=6%) % 80         *
               
Rabbit nutritive values Unit Avg SD Min Max Nb  
DE rabbit MJ/kg DM 15.7       1 *
MEn rabbit MJ/kg DM 15.1         *
Energy digestibility, rabbit % 83.3       1 *
Nitrogen digestibility, rabbit % 78.9       1 *

The asterisk * indicates that the average value was obtained by an equation.

References

Adeola, 2003; AFZ, 2025; CIRAD, 1991; De Silva et al., 1990; Eeckhout et al., 1994; Jondreville et al., 2000; Krishna, 1985; Lekule et al., 1990; Maertens et al., 1985; Mariscal Landin, 1992; Noblet et al., 2000; Rajaguru et al., 1985; Tiwari et al., 2006

Last updated on 30/10/2026 16:22:53

Main analysis Unit Avg SD Min Max Nb  
Dry matter % as fed 88.1 1.3 85 95.7 353  
Crude protein % DM 17.9 1.2 13.5 21.2 348  
Crude fibre % DM 8.2 1.6 3.4 11.4 140  
Neutral detergent fibre % DM 36.2 6.1 22.7 50.3 80 *
Acid detergent fibre % DM 10.6 2.3 5.3 15.2 88 *
Lignin % DM 3.1 0.8 1.4 4.3 67 *
Ether extract % DM 4.3 0.9 0.9 6.1 347  
Ash % DM 4.4 0.8 2.3 6.1 103  
Insoluble ash % DM 0.09 0.04 0.02 0.2 27  
Starch (polarimetry) % DM 21.4 0.9 19.2 22.9 116  
Starch (enzymatic) % DM 17.8         *
Total sugars % DM 7.9 1.9 2.4 8.2 10 *
Gross energy MJ/kg DM 19.2 0.4 18.4 20.3 30 *
               
Amino acids Unit Avg SD Min Max Nb  
Alanine g/16g N 4.5 0.4 3.4 5.2 47 *
Arginine g/16g N 6.4 0.6 4.9 7.2 49 *
Aspartic acid g/16g N 6.6 0.4 5.5 8.1 46 *
Cystine g/16g N 2 0.1 1.6 2.4 41 *
Glutamic acid g/16g N 20.2 2.8 14.4 26.7 47 *
Glycine g/16g N 4.8 0.5 3.5 5.8 50 *
Histidine g/16g N 2.6 0.2 2 3 48 *
Isoleucine g/16g N 3.2 0.2 2.8 3.8 53 *
Leucine g/16g N 6.1 0.4 5.2 7 53 *
Lysine g/16g N 4 0.5 2.2 4.9 56 *
Methionine g/16g N 1.5 0.2 0.8 2 50 *
Methionine+cystine g/16g N 3.6 0.3 3.1 4.3 41 *
Phenylalanine g/16g N 4 0.2 3.4 4.4 48 *
Phenylalanine+tyrosine g/16g N 6.6 0.4 5.7 7.3 29 *
Proline g/16g N 6.7 0.8 5.1 8.7 34 *
Serine g/16g N 4.3 0.3 3.1 5 47 *
Threonine g/16g N 3.2 0.3 2.6 4 53 *
Tryptophan g/16g N 1.3 0.2 1.1 1.8 33 *
Tyrosine g/16g N 2.6 0.3 2.1 3.5 37 *
Valine g/16g N 4.5 0.3 3.8 5.1 49 *
               
Fatty acids Unit Avg SD Min Max Nb  
Myristic acid C14:0 % fatty acids 0.1   0 0.3 4  
Palmitic acid C16:0 % fatty acids 17.8 0.9 15.5 18.3 9  
Palmitoleic acid C16:1 % fatty acids 0.4          
Stearic acid C18:0 % fatty acids 0.8 0.07 0.7 0.9 8  
Oleic acid C18:1 % fatty acids 15.2 2.7 11.3 20.4 11  
Linoleic acid C18:2 % fatty acids 56.4          
Linolenic acid C18:3 % fatty acids 5.9 1 5.4 8.5 9  
               
Minerals Unit Avg SD Min Max Nb  
Calcium g/kg DM 1.2 0.4 0.2 2.4 68 *
Phosphorus g/kg DM 9 2.2 5.2 16.8 77 *
Potassium g/kg DM 11.3 1.8 8.1 16.9 40 *
Sodium g/kg DM 0.09 0.01 0.09 0.12 11  
Chlorine g/kg DM 1.1          
Magnesium g/kg DM 2.6 0.8 1.2 4.8 27  
Sulfur g/kg DM 2.3          
Manganese mg/kg DM 99 35 58 169 13  
Zinc mg/kg DM 95 18 72 139 13  
Copper mg/kg DM 15 2 12 17 11  
Iron mg/kg DM 69   65 72 2  
Selenium mg/kg DM 0.2          
               
Pig nutritive values Unit Avg SD Min Max Nb  
Energy digestibility, growing pig % 65.7   56.4 78 4 *
DE growing pig MJ/kg DM 12.6         *
MEn growing pig MJ/kg DM 12       1 *
NE growing pig MJ/kg DM 8.3         *
Nitrogen digestibility, growing pig % 70.3   69 79.4 3 *
               
Poultry nutritive values Unit Avg SD Min Max Nb  
AMEn cockerel MJ/kg DM 9.8         *
AMEn broiler MJ/kg DM 9.4         *
               
Ruminants nutritive values Unit Avg SD Min Max Nb  
OM digestibility, ruminants % 74.6 5.2 71.6 87.2 21 *
Energy digestibility, ruminants % 72.9 5.7 67.1 84.7 20 *
ME ruminants MJ/kg DM 11.4         *
Nitrogen digestibility, ruminants % 68.6         *
Nitrogen degradability (effective, k=6%) % 76 8 68 87 5 *
Nitrogen degradability (effective, k=4%) % 80       1 *
a (N) % 35   32 56 2  
b (N) % 57   39 83 3  
c (N) h-1 0.155   0.134 0.182 3  
Dry matter degradability (effective, k=6%) % 72 3 70 77 5 *
Dry matter degradability (effective, k=4%) % 75       1 *
a (DM) % 46          
b (DM) % 38          
c (DM) h-1 0.13 0.057 0.004 0.145 5  
               
Rabbit nutritive values Unit Avg SD Min Max Nb  
DE rabbit MJ/kg DM 13.9         *
MEn rabbit MJ/kg DM 13.3         *
Energy digestibility, rabbit % 72.4       1 *
Nitrogen digestibility, rabbit % 59.8         *

The asterisk * indicates that the average value was obtained by an equation.

References

ADAS, 1989; Adedokun et al., 2015; AFZ, 2025; Allan et al., 2000; Anderson et al., 1991; Anon., 2001; Aregheore, 2000; Aufrère et al., 1991; Aureli et al., 2015; Bach Knudsen, 1997; Barrows et al., 2015; Bryden et al., 2009; CIRAD, 1991; CIRAD, 2008; Clark et al., 1997; Cottyn et al., 1978; De Boever et al., 1994; Dei et al., 2008; Devendra et al., 1970; Dewar, 1967; Eeckhout et al., 1994; El-Haroun et al., 2007; Erdman et al., 1987; Fernandez Carmona et al., 1996; Getachew et al., 2004; Hajen et al., 1993; Han et al., 1976; Hanczakowski et al., 1979; Hepburn et al., 1960; Holtshausen et al., 2011; Huang et al., 2007; Jongbloed et al., 1990; Kendall et al., 1982; Kiiskinen, 1992; Lechevestrier, 1992; Leeson et al., 1974; Lekule et al., 1990; Macgregor et al., 1978; Maertens et al., 1985; Marcondes et al., 2009; Mariscal Landin, 1992; Masoero et al., 1994; May et al., 1971; McNab et al., 1988; Morgan et al., 1975; Morse et al., 1992; Nengas et al., 1995; Oliveira et al., 2007; Petit, 1992; Pozy et al., 1996; Qiao ShiYan et al., 2004; Rogerson, 1956; San Juan et al., 1993; Sauer et al., 1989; Smolders et al., 1990; Storey et al., 1982; Swanek et al., 2001; Tahir et al., 2012; Tamminga et al., 1990; Tesfaye et al., 2013; Wainman et al., 1979

Last updated on 30/10/2026 16:24:37

Main analysis Unit Avg SD Min Max Nb  
Dry matter % as fed 88 1.1 83.8 96.7 1497  
Crude protein % DM 18.1 1 13.5 21.3 1508  
Crude fibre % DM 8 1.1 3.4 11.7 618  
Neutral detergent fibre % DM 35.5 5.9 22.7 50.3 104 *
Acid detergent fibre % DM 10.4 2.2 5.3 15.2 112 *
Lignin % DM 3 0.7 1.4 4.3 136 *
Ether extract % DM 4.6 0.6 0.9 6 373  
Ash % DM 4.3 0.5 2.3 6.4 505  
Insoluble ash % DM 0.09 0.04 0.01 0.2 29  
Starch (polarimetry) % DM 28.6 2.9 23 34 1261  
Starch (enzymatic) % DM 24.5         *
Total sugars % DM 8 2.2 2.4 12 70 *
Gross energy MJ/kg DM 19.3 0.3 18.4 20.3 33 *
               
Amino acids Unit Avg SD Min Max Nb  
Alanine g/16g N 4.5 0.4 3.4 5.2 52 *
Arginine g/16g N 6.4 0.5 4.9 7.2 54 *
Aspartic acid g/16g N 6.7 0.4 5.5 8.1 51 *
Cystine g/16g N 2 0.2 1.6 2.6 47 *
Glutamic acid g/16g N 20.1 2.7 14.4 26.7 51 *
Glycine g/16g N 4.8 0.4 3.5 5.8 55 *
Histidine g/16g N 2.6 0.2 2 3.1 53 *
Isoleucine g/16g N 3.2 0.2 2.8 3.8 58 *
Leucine g/16g N 6.1 0.4 5.2 7 58 *
Lysine g/16g N 4 0.5 2.2 4.9 63 *
Methionine g/16g N 1.5 0.2 0.8 2 56 *
Methionine+cystine g/16g N 3.6 0.3 3.1 4.4 46 *
Phenylalanine g/16g N 4 0.2 3.4 4.4 53 *
Phenylalanine+tyrosine g/16g N 6.5 0.4 5.7 7.3 32 *
Proline g/16g N 6.6 0.7 5.1 8.7 37 *
Serine g/16g N 4.3 0.3 3.1 5 52 *
Threonine g/16g N 3.2 0.3 2.6 4 58 *
Tryptophan g/16g N 1.3 0.2 1.1 1.8 35 *
Tyrosine g/16g N 2.6 0.3 2.1 3.5 40 *
Valine g/16g N 4.5 0.3 3.8 5.1 54 *
               
Fatty acids Unit Avg SD Min Max Nb  
Myristic acid C14:0 % fatty acids 0.1   0 0.3 4  
Palmitic acid C16:0 % fatty acids 17.8 0.9 15.5 18.3 9  
Palmitoleic acid C16:1 % fatty acids 0.4          
Stearic acid C18:0 % fatty acids 0.8 0.07 0.7 0.9 8  
Oleic acid C18:1 % fatty acids 15.2 2.7 11.3 20.4 11  
Linoleic acid C18:2 % fatty acids 56.4          
Linolenic acid C18:3 % fatty acids 5.9 1 5.4 8.5 9  
               
Minerals Unit Avg SD Min Max Nb  
Calcium g/kg DM 1.2 0.4 0.2 2.4 168 *
Phosphorus g/kg DM 8.8 1.8 5.2 16.8 182 *
Potassium g/kg DM 11 1.7 8.1 16.9 46 *
Sodium g/kg DM 0.06 0.04 0.01 0.2 55  
Chlorine g/kg DM 1   0.8 1.2 3  
Magnesium g/kg DM 3.3 0.8 1.2 4.8 31  
Sulfur g/kg DM 1.9 0.3 1.5 2.3 5  
Manganese mg/kg DM 109 31 58 169 21  
Zinc mg/kg DM 99 19 72 139 16  
Copper mg/kg DM 15 2 11 17 14  
Iron mg/kg DM 143 71 65 215 5  
Selenium mg/kg DM 0.2       1  
               
Pig nutritive values Unit Avg SD Min Max Nb  
Energy digestibility, growing pig % 66.3   56.4 78 4 *
DE growing pig MJ/kg DM 12.8         *
MEn growing pig MJ/kg DM 12.2       1 *
NE growing pig MJ/kg DM 8.7         *
Nitrogen digestibility, growing pig % 70.7   69 79.4 3 *
               
Poultry nutritive values Unit Avg SD Min Max Nb  
AMEn cockerel MJ/kg DM 9.9         *
AMEn broiler MJ/kg DM 9.5         *
               
Ruminants nutritive values Unit Avg SD Min Max Nb  
OM digestibility, ruminants % 74.9 5.2 71.6 87.2 21 *
Energy digestibility, ruminants % 73.2 5.7 67.1 84.7 20 *
ME ruminants MJ/kg DM 11.6         *
Nitrogen digestibility, ruminants % 68.8         *
Nitrogen degradability (effective, k=6%) % 76 8 68 87 6 *
Nitrogen degradability (effective, k=4%) % 80       1 *
a (N) % 35   32 56 2  
b (N) % 57   39 83 3  
c (N) h-1 0.155   0.134 0.182 3  
Dry matter degradability (effective, k=6%) % 64         *
Dry matter degradability (effective, k=4%) % 68       1 *
a (DM) % 44   27 74 3  
b (DM) % 38 8 21 46 6  
c (DM) h-1 0.065 0.052 0.004 0.145 6  
               
Rabbit nutritive values Unit Avg SD Min Max Nb  
DE rabbit MJ/kg DM 14         *
MEn rabbit MJ/kg DM 13.4         *
Energy digestibility, rabbit % 72.6       1 *
Nitrogen digestibility, rabbit % 60.9         *

The asterisk * indicates that the average value was obtained by an equation.

References

ADAS, 1988; ADAS, 1989; Adedokun et al., 2015; AFZ, 2025; Allan et al., 2000; Anderson et al., 1991; Anon., 2001; Aufrère et al., 1991; Aureli et al., 2015; Bach Knudsen, 1997; Barrows et al., 2015; Batajoo et al., 1998; Bryden et al., 2009; CIRAD, 1991; CIRAD, 2008; Clark et al., 1997; Cowan et al., 1998; De Boever et al., 1994; Dei et al., 2008; Dewar, 1967; Eeckhout et al., 1994; El-Haroun et al., 2007; Erdman et al., 1987; Fernandez Carmona et al., 1996; Getachew et al., 2004; Hajen et al., 1993; Han et al., 1976; Hanczakowski et al., 1979; Hepburn et al., 1960; Holtshausen et al., 2011; Huang et al., 2007; Jondreville et al., 2000; Jongbloed et al., 1990; Kendall et al., 1982; Kiiskinen, 1992; Lechevestrier, 1992; Lekule et al., 1990; Macgregor et al., 1978; Maertens et al., 1985; Marcondes et al., 2009; Mariscal Landin, 1992; May et al., 1971; Morgan et al., 1975; Morse et al., 1992; Oliveira et al., 2007; Petit, 1992; Pozy et al., 1996; Qiao ShiYan et al., 2004; Rogerson, 1956; San Juan et al., 1993; Sauer et al., 1989; Smolders et al., 1990; Storey et al., 1982; Swanek et al., 2001; Tahir et al., 2012; Tamminga et al., 1990; Tesfaye et al., 2013; Wainman et al., 1979

Last updated on 30/10/2026 16:25:52

Main analysis Unit Avg SD Min Max Nb  
Dry matter % as fed 87.9 1.1 84.3 92.9 793  
Crude protein % DM 17.6 1.2 10.8 21.3 792  
Crude fibre % DM 6.8 1.3 3.4 10.6 322  
Neutral detergent fibre % DM 30.7 6.6 20.8 50.3 97 *
Acid detergent fibre % DM 8.8 2.3 5.3 15.2 105 *
Lignin % DM 2.6 0.9 1.3 4.3 97 *
Ether extract % DM 4.2 0.7 0.9 5.6 188  
Ash % DM 3.8 0.5 2.3 5.6 238  
Insoluble ash % DM 0.09 0.04 0.02 0.2 27  
Starch (polarimetry) % DM 38.5 3.1 34 45.2 556  
Starch (enzymatic) % DM 33.7 9.7 25.1 57.5 14 *
Total sugars % DM 7.7 2.1 2.4 10.1 33 *
Gross energy MJ/kg DM 19.2 0.4 18.4 20.3 33 *
               
Amino acids Unit Avg SD Min Max Nb  
Alanine g/16g N 4.4 0.4 3.4 5.2 51 *
Arginine g/16g N 6.3 0.5 4.9 7.2 53 *
Aspartic acid g/16g N 6.6 0.4 5.5 8.1 50 *
Cystine g/16g N 2 0.1 1.6 2.4 45 *
Glutamic acid g/16g N 20.2 2.7 14.4 26.7 51 *
Glycine g/16g N 4.8 0.4 3.5 5.8 54 *
Histidine g/16g N 2.7 0.2 2 3 52 *
Isoleucine g/16g N 3.2 0.2 2.8 3.8 57 *
Leucine g/16g N 6.1 0.4 5.2 7 57 *
Lysine g/16g N 4 0.5 2.2 4.9 61 *
Methionine g/16g N 1.5 0.2 0.8 2 54 *
Methionine+cystine g/16g N 3.5 0.3 3.1 4.3 45 *
Phenylalanine g/16g N 4 0.2 3.4 4.4 52 *
Phenylalanine+tyrosine g/16g N 6.6 0.4 5.7 7.3 30 *
Proline g/16g N 6.7 0.7 5.1 8.7 35 *
Serine g/16g N 4.3 0.3 3.1 5 51 *
Threonine g/16g N 3.2 0.3 2.6 4 57 *
Tryptophan g/16g N 1.3 0.2 1.1 1.8 36 *
Tyrosine g/16g N 2.6 0.3 2.1 3.5 38 *
Valine g/16g N 4.5 0.3 3.8 5.1 53 *
               
Fatty acids Unit Avg SD Min Max Nb  
Myristic acid C14:0 % fatty acids 0.1   0 0.3 4  
Palmitic acid C16:0 % fatty acids 17.8 0.9 15.5 18.3 9  
Palmitoleic acid C16:1 % fatty acids 0.4          
Stearic acid C18:0 % fatty acids 0.8 0.07 0.7 0.9 8  
Oleic acid C18:1 % fatty acids 15.2 2.7 11.3 20.4 11  
Linoleic acid C18:2 % fatty acids 56.4       1  
Linolenic acid C18:3 % fatty acids 5.9 1 5.4 8.5 9  
               
Minerals Unit Avg SD Min Max Nb  
Calcium g/kg DM 1.1 0.4 0.2 2.4 103 *
Phosphorus g/kg DM 7.9 2.2 5.2 16.8 117 *
Potassium g/kg DM 10.1 1.8 8.1 16.9 40 *
Sodium g/kg DM 0.06 0.04 0.01 0.12 22  
Chlorine g/kg DM 1.1          
Magnesium g/kg DM 3.3 0.8 1.2 4.8 27  
Sulfur g/kg DM 2.3   2.3 2.3 2  
Manganese mg/kg DM 99 31 58 169 16  
Zinc mg/kg DM 95 18 72 139 13  
Copper mg/kg DM 15 2 12 17 11  
Iron mg/kg DM 69   65 72 2  
Selenium mg/kg DM 0.2       1  
               
Pig nutritive values Unit Avg SD Min Max Nb  
Energy digestibility, growing pig % 71.1 7.2 56.4 78 6 *
DE growing pig MJ/kg DM 13.6         *
MEn growing pig MJ/kg DM 13.1       1 *
NE growing pig MJ/kg DM 9.6         *
Nitrogen digestibility, growing pig % 73.6   69 79.4 4 *
               
Poultry nutritive values Unit Avg SD Min Max Nb  
AMEn cockerel MJ/kg DM 10.8         *
AMEn broiler MJ/kg DM 10.3         *
               
Ruminants nutritive values Unit Avg SD Min Max Nb  
OM digestibility, ruminants % 77 5.2 71.6 87.2 21 *
Energy digestibility, ruminants % 75.3 5.7 67.1 84.7 20 *
ME ruminants MJ/kg DM 11.9         *
Nitrogen digestibility, ruminants % 69.1         *
Nitrogen degradability (effective, k=6%) % 76 8 68 87 5 *
Nitrogen degradability (effective, k=4%) % 80       1 *
a (N) % 35   32 56 2  
b (N) % 57   39 83 3  
c (N) h-1 0.155   0.134 0.182 3  
Dry matter degradability (effective, k=6%) % 64         *
Dry matter degradability (effective, k=4%) % 68       1 *
a (DM) % 44          
b (DM) % 38          
c (DM) h-1 0.065 0.057 0.004 0.145 5  
               
Rabbit nutritive values Unit Avg SD Min Max Nb  
DE rabbit MJ/kg DM 14.3         *
MEn rabbit MJ/kg DM 13.7         *
Energy digestibility, rabbit % 74.5       1 *
Nitrogen digestibility, rabbit % 65.1         *

The asterisk * indicates that the average value was obtained by an equation.

References

ADAS, 1989; Adedokun et al., 2015; AFZ, 2025; Allan et al., 2000; Anderson et al., 1991; Anon., 2001; Aufrère et al., 1991; Aureli et al., 2015; Bach Knudsen, 1997; Barrows et al., 2015; Bryden et al., 2009; Castaing et al., 1995; CIRAD, 1991; CIRAD, 2008; Clark et al., 1997; De Boever et al., 1994; Dei et al., 2008; Dewar, 1967; Eeckhout et al., 1994; El-Haroun et al., 2007; Erdman et al., 1987; Fernandez Carmona et al., 1996; Getachew et al., 2004; Guillaume, 1978; Hajen et al., 1993; Han et al., 1976; Hanczakowski et al., 1979; Hepburn et al., 1960; Holtshausen et al., 2011; Huang et al., 2007; Jondreville et al., 2000; Jongbloed et al., 1990; Kendall et al., 1982; Kiiskinen, 1992; Lechevestrier, 1992; Lekule et al., 1990; Macgregor et al., 1978; Maertens et al., 1985; Marcondes et al., 2009; Mariscal Landin, 1992; May et al., 1971; Morgan et al., 1975; Morse et al., 1992; Noblet et al., 2000; Oliveira et al., 2007; Petit, 1992; Pozy et al., 1996; Qiao ShiYan et al., 2004; Rogerson, 1956; San Juan et al., 1993; Sauer et al., 1989; Shi et al., 1993; Smolders et al., 1990; Storey et al., 1982; Swanek et al., 2001; Tahir et al., 2012; Tamminga et al., 1990; Tesfaye et al., 2013; Wainman et al., 1979

Last updated on 30/10/2026 16:28:09

Main analysis Unit Avg SD Min Max Nb  
Dry matter % as fed 87.8 1.2 84 92.9 370  
Crude protein % DM 17.1 1.5 13.1 21.2 362  
Crude fibre % DM 6.5 1.9 3 10.4 137  
Neutral detergent fibre % DM 29.4 6.7 20.6 50.3 83 *
Acid detergent fibre % DM 8.4 2.4 5.2 15.2 91 *
Lignin % DM 2.5 1 0.9 4.3 74 *
Ether extract % DM 3.9 1 0.9 6.1 455  
Ash % DM 3.6 0.9 1.9 5.6 99  
Insoluble ash % DM 0.09 0.04 0.02 0.2 28  
Starch (polarimetry) % DM 51.1 5 45.5 67.6 133  
Starch (enzymatic) % DM 45.4 9.7 25.1 57.5 14 *
Total sugars % DM 7.4 2.3 2.4 8.9 8 *
Gross energy MJ/kg DM 19.1 0.4 18.4 20.3 30 *
               
Amino acids Unit Avg SD Min Max Nb  
Alanine g/16g N 4.4 0.4 3.4 5.2 47 *
Arginine g/16g N 6.2 0.6 4.9 7.2 49 *
Aspartic acid g/16g N 6.5 0.4 5.5 8.1 46 *
Cystine g/16g N 2 0.1 1.6 2.4 41 *
Glutamic acid g/16g N 20.2 2.8 14.4 26.7 47 *
Glycine g/16g N 4.8 0.5 3.5 5.8 50 *
Histidine g/16g N 2.7 0.2 2 3 48 *
Isoleucine g/16g N 3.2 0.2 2.8 3.8 53 *
Leucine g/16g N 6.1 0.4 5.2 7 53 *
Lysine g/16g N 3.9 0.5 2.2 4.9 59 *
Methionine g/16g N 1.5 0.2 0.8 2 50 *
Methionine+cystine g/16g N 3.5 0.3 3.1 4.3 41 *
Phenylalanine g/16g N 4 0.2 3.4 4.4 48 *
Phenylalanine+tyrosine g/16g N 6.7 0.4 5.7 7.3 29 *
Proline g/16g N 6.7 0.8 5.1 8.7 34 *
Serine g/16g N 4.3 0.3 3.1 5 47 *
Threonine g/16g N 3.2 0.3 2.6 4 53 *
Tryptophan g/16g N 1.3 0.2 1.1 1.8 33 *
Tyrosine g/16g N 2.7 0.3 2.1 3.5 37 *
Valine g/16g N 4.5 0.3 3.8 5.1 49 *
               
Fatty acids Unit Avg SD Min Max Nb  
Myristic acid C14:0 % fatty acids 0.1   0 0.3 4  
Palmitic acid C16:0 % fatty acids 17.8 0.9 15.5 18.3 9  
Palmitoleic acid C16:1 % fatty acids 0.4          
Stearic acid C18:0 % fatty acids 0.8 0.07 0.7 0.9 8  
Oleic acid C18:1 % fatty acids 15.2 2.7 11.3 20.4 11  
Linoleic acid C18:2 % fatty acids 56.4          
Linolenic acid C18:3 % fatty acids 5.9 1 5.4 8.5 9  
               
Minerals Unit Avg SD Min Max Nb  
Calcium g/kg DM 1.1 0.4 0.2 2.4 71 *
Phosphorus g/kg DM 7.5 2.6 4.4 16.8 80 *
Potassium g/kg DM 9.7 1.8 8.1 16.9 40 *
Sodium g/kg DM 0.1 0.02 0.08 0.17 13  
Chlorine g/kg DM 1.1          
Magnesium g/kg DM 3.3 0.8 1.2 4.8 27  
Sulfur g/kg DM 1.9          
Manganese mg/kg DM 99 35 58 169 13  
Zinc mg/kg DM 95 18 72 139 13  
Copper mg/kg DM 15 2 12 17 11  
Iron mg/kg DM 69   65 72 2  
Selenium mg/kg DM 0.2          
               
Pig nutritive values Unit Avg SD Min Max Nb  
Energy digestibility, growing pig % 72.2   56.4 78 4 *
DE growing pig MJ/kg DM 13.8         *
MEn growing pig MJ/kg DM 13.3       1 *
NE growing pig MJ/kg DM 10.2         *
Nitrogen digestibility, growing pig % 74.3   69 79.4 3 *
               
Poultry nutritive values Unit Avg SD Min Max Nb  
AMEn cockerel MJ/kg DM 11         *
AMEn broiler MJ/kg DM 10.5         *
               
Ruminants nutritive values Unit Avg SD Min Max Nb  
OM digestibility, ruminants % 77.5 5.2 71.6 87.2 21 *
Energy digestibility, ruminants % 75.7 5.7 67.1 84.7 20 *
ME ruminants MJ/kg DM 11.9         *
Nitrogen digestibility, ruminants % 68.9         *
Nitrogen degradability (effective, k=6%) % 76 8 68 87 5 *
Nitrogen degradability (effective, k=4%) % 80       1 *
a (N) % 35   32 56 2  
b (N) % 57   39 83 3  
c (N) h-1 0.155   0.134 0.182 3  
Dry matter degradability (effective, k=6%) % 64         *
Dry matter degradability (effective, k=4%) % 68       1 *
a (DM) % 44          
b (DM) % 38          
c (DM) h-1 0.065 0.057 0.004 0.145 5  
               
Rabbit nutritive values Unit Avg SD Min Max Nb  
DE rabbit MJ/kg DM 14.4         *
MEn rabbit MJ/kg DM 13.7         *
Energy digestibility, rabbit % 75.1       1 *
Nitrogen digestibility, rabbit % 66         *

The asterisk * indicates that the average value was obtained by an equation.

References

ADAS, 1989; Adedokun et al., 2015; AFZ, 2025; Allan et al., 2000; Anderson et al., 1991; Anon., 2001; Aregheore, 2000; Aufrère et al., 1991; Aureli et al., 2015; Bach Knudsen, 1997; Barrows et al., 2015; Bryden et al., 2009; CIRAD, 1991; Clark et al., 1997; Cottyn et al., 1978; De Boever et al., 1994; Dei et al., 2008; Devendra et al., 1970; Dewar, 1967; Eeckhout et al., 1994; El-Haroun et al., 2007; Erdman et al., 1987; Fernandez Carmona et al., 1996; Getachew et al., 2004; Hajen et al., 1993; Han et al., 1976; Hanczakowski et al., 1979; Hepburn et al., 1960; Holtshausen et al., 2011; Huang et al., 2007; Jongbloed et al., 1990; Kendall et al., 1982; Kiiskinen, 1992; Lechevestrier, 1992; Leeson et al., 1974; Lekule et al., 1990; Macgregor et al., 1978; Maertens et al., 1985; Marcondes et al., 2009; Mariscal Landin, 1992; Masoero et al., 1994; May et al., 1971; McNab et al., 1988; Morgan et al., 1975; Morse et al., 1992; Nengas et al., 1995; Oliveira et al., 2007; Petit, 1992; Pozy et al., 1996; Qiao ShiYan et al., 2004; Rogerson, 1956; San Juan et al., 1993; Sauer et al., 1989; Smolders et al., 1990; Storey et al., 1982; Swanek et al., 2001; Tahir et al., 2012; Tamminga et al., 1990; Tesfaye et al., 2013; Wainman et al., 1979

Last updated on 30/10/2026 16:29:02

References
References 
Datasheet citation 

Heuzé V., Tran G., 2026. Wheat shorts, wheat middlings and feed flour. Feedipedia, a programme by INRAE, CIRAD, AFZ and FAO. https://www.feedipedia.org/node/727 Last updated on November 2, 2026, 9:52