Maize silage has become the predominant forage fed to ruminant livestock in many parts of Europe, Australia, New Zealand, North America and India, offered alone or as a complement to grass silage. Its main advantages are a high starch content, a high protein efficiency ratio and a relatively high content of digestible energy and nutrients (Fitzgerald et al., 1998; INRA, 2007; Keady et al., 2013; Malisetty et al., 2013). However, maize silage has a low protein content, which declines as maturity increases (INRA, 2007; Keady et al., 2013). Diets including maize silage are usually supplemented with sources of energy and protein, particularly for high-producing livestock.
Digestibility and energy values
The OM digestibility of maize silage ranges from 62% to 76%, depending on the growing conditions and on the stage of growth at harvest (Barrière et al., 2003; INRA, 2007). OM digestibility was also shown to vary between different maize hybrids depending on their earliness. OM digestibility was comprised between 72% for very early and 67% for very late hybrids (Michalet-Doreau et al., 1999). The amount and the composition of whole plant cell walls, especially their lignification, influence both digestibility and voluntary intake when maize is given as silage (Barrière et al., 2003). Starch is degraded slowly in the rumen, and differences among genotypes exist. Starch degradability also depends on plant maturity, decreasing from more than 80% to less than 50% when whole plant DM increased from 25% to 40% (Michalet-Doreau et al., 1999). In the case of baby corn husks, in vitro OM digestibility (83 vs. 79%) and ME content (10.2 vs. 9.2 MJ/kg DM) were higher for fresh baby corn husks, compared to baby corn husk silage (Bakshi et al., 2012).
Intake and palatability
Voluntary intake of maize silage varies greatly. Well-preserved silages can be ingested at levels similar to the fresh or dry maize forages, but intake levels decrease with preservation quality. Badly preserved silages may have a low palatability. Acidity has a negative effect on maize silage intake in cattle, as it is generally negatively linked to DM content, and cattle, unlike sheep, are very sensitive to this. The low protein level of maize silage, when not supplemented, can negatively affect intake (Dulphy et al., 1996). In India, baby corn husk silage offered as a total mixed ration, with wheat straw and a concentrate mixture, to male buffalo calves, resulted in higher nutrient digestibility, voluntary intake and N retention than for fresh baby corn husks (Bakshi et al., 2012).
Dairy cows
Effect of hybrid types
The nutritional quality of maize silage can be improved through hybrid selection, benefiting dairy farmers through improved feed efficiency and milk yield. A meta-analysis of 48 papers published between 1995 and 2014 showed that intake, digestibility, rumen fermentation and lactation performance by dairy cows were affected by the type of hybrid ensiled (see Table 1 below). Hybrids selected for greater fibre digestibility increased milk and protein yield through a higher DM intake, but failed to improve feed efficiency and reduced starch digestibility. Lactation performance was little improved by non-GM hybrids and not improved at all by GM hybrids (Ferrareto et al., 2015).
Table 1: Effect of maize silage hybrids on lactation performance of dairy cows (range of least square means; SEM in brackets), adapted from Ferrareto et al., 2015.
Treatment |
DMI, kd/d |
Milk, kg/d |
Milk fat, kg/d |
Milk protein, kg/d |
Maize silage hybrids with differing stalk characteristics |
23.7-24.9 (0.4) |
37.2-38.7 (0.8) |
1.34-1.37 (0.03) |
1.13-1.18
(0.02 |
Maize silage hybrids with differing grain characteristics |
23.6-24.1 (1.1) |
36.1-37.7 (1.9) |
1.20-1.33 (0.08) |
1.11-1.16
(0.06) |
Genetically modified maize silage hybrids |
24.3-24.4 (0.7) |
34.8-35.0 (1.3) |
1.25-1.26 (0.04) |
1.11
(0.03) |
Use of maize silage in grass silage based diets
A review of 34 comparisons in which grass silage was partially or totally replaced with maize silage, in diets for lactating cows, showed that maize silage significantly increased forage DM intake (+1.5 kg/d) and milk yield (+1.4 kg/d milk, +0.6 g/kg fat, and +0.86g/kg protein). Including maize silage in grass silage based diets could potentially reduce the concentrates needed by up to 5 kg/cow/d (Keady et al., 2013).
From a series of experiments conducted in Ireland, it was shown that the inclusion of maize silage in diets with grass silage increased forage intake. However, the benefit in milk production depended on the maturity (starch content) and digestibility (greenness) of the crop when harvested (see Table 2 below). In particular, partially replacing a high quality grass silage with good quality maize silage (starch 20-25% of DM) increased forage intake, milk yield and milk protein concentration without affecting the efficiency of milk production. Moreover, the inclusion of 33% maize silage in the forage allocation achieved most of the increase in milk yield, but a higher inclusion level (67%) was required to maximize the yields of milk, fat, and protein. In practice a 50:50 mixture of maize silage and grass silage seemed to be near the optimum for milk production (Fitzgerald et al., 1998).
Table 2: Effect of inclusion of maize silage on lactation performance of dairy cows (means; SEM in brackets), adapted from Fitzgerald et al., 1998.
Treatment |
|
Silage intake, kg/d |
Milk, kg/d |
Milk fat, kg/d |
Milk protein, kg/d |
% of inclusion of maize silage with grass silage |
0% |
8.8 (0.35) |
21.4 (0.48) |
0.81 (0.02) |
0.65 (0.01) |
|
33% |
9.7 (0.35) |
23.0 (0.48) |
0.84 (0.02) |
0.71 (0.01) |
|
67% |
10.4 (0.35) |
23.1 (0.48) |
0.86 (0.02) |
0.73 (0.01) |
|
100% |
10.7 (0.35) |
22.7 (0.48) |
0.84 (0.02) |
0.70 (0.01) |
High quality grass silagea |
8.9 (0.32) |
18.6 (0.70) |
0.85 (0.03) |
0.66 (0.02) |
Moderate starch maize silage (242 g/kg DM)
+ grass silage (60:40)a |
11.1 (0.32) |
20.5 (0.70) |
0.91 (0.03) |
0.73 (0.02) |
High starch maize silage
(293 g/kg DM) + grass silage (60:40)a |
11.7 (0.32) |
19.5 (0.70) |
0.88 (0.03) |
0.71 (0.02) |
a The animals were supplemented with 4-8 kg/d concentrate.
Growing cattle
A review of 9 studies in which grass silage was partially or totally replaced by maize silage showed that maize silage significantly increased forage DM intake (+1.5 kg/d) and growth of beef cattle (+0.23 kg/d live-weight gain; +0.11 kg/d carcass gain; +12 kg carcass weight). When offered ad libitum as the sole forage, daily carcass gain was increased by 31% due to a combination of increased metabolizable energy intake and improved efficiency of metabolizable energy utilization. Finally, a potential concentrate sparing effect of up to 2.4 kg/d/beef cattle was reported when maize silage was included in the diet. The effect depended on the level of maize included in the diet and on the crop maturity at harvest (Keady et al., 2013).
Sheep
In many sheep-producing areas of Europe, animals are housed during the winter feeding period and offered ensiled forage. Few studies have been undertaken to evaluate the effects of maize silage on the performance of pregnant ewes or finishing lambs. However, its low protein content may impact its ability to meet the protein requirements of animals with high requirements. A review of studies conducted in the UK concluded that maize silage can replace high-feed value grass silage in the diet of ewes in mid and late-pregnancy (see Table 3 below). Maize silage, whilst low in protein, can be offered as the sole forage without protein supplementation until late pregnancy (final 6 to 7 weeks). Increasing the maturity of maize at harvest tended to increase lamb weaning weight by 1 kg (Keady et al., 2013).
Table 3: Effect of maize silage maturity and grass silage feed value on ewe and lamb performance (range of least square means; SEM in brackets), adapted from Keady et al., 2013.
Treatmenta |
Ewe weight post lambing, kg |
Lamb weight at birth, kg |
Lamb weight at weaning, kg |
Carcass weight, kg |
Maize silage with low DM content |
63.0-68.6 (2.2) |
4.6-4.9 (0.2) |
32.8-33.4 (1.1) |
18.2-18.9 (0.4) |
Maize silage with high DM content |
68.2-76.6 (2.2) |
4.7-5.3 (0.2) |
34.1-34.3 (1.1) |
18.7-19.1 (0.4) |
Grass silage with low feed-value |
61.2-61.6 (2.2) |
4.6 (0.2) |
32.1-33.6 (1.1) |
18.8-19.1 (0.4) |
Grass silage with high feed-value |
70.4-73.6 (2.2) |
4.8-5.1 (0.2) |
34.0-35.0 (1.1) |
19.2-19.6 (0.4) |
a The animals were supplemented with 0 or 200 g/d soybean meal, and/or 5 to 25 kg concentrate (total concentrate during late pregnancy).
Regarding finishing lambs, increasing maize maturity at the time of ensiling (and therefore decreasing starch content) improved carcass gain. Maize silage replaced medium and high feed-value grass silage in the diet, resulting in equivalent or improved daily carcass gain, respectively (see Table 4 below). Replacing medium feed-value grass silage with maize silage resulted in a potential concentrate sparing effect of up to 0.48 kg/lamb/d (Keady et al., 2013).
Table 4: Effect of forage type on lamb performance (range of means), adapted from Keady et al., 2013.
Treatmenta |
Total DM intake, kg/d |
Carcass weight, kg |
Carcass gain, g/d |
Maize silage |
1.1-1.3 |
20.2-23.3 |
59-107 |
Medium feed-value grass silage |
0.9-1.2 |
17.3-22.8 |
14-98 |
High feed-value grass silage |
1.1-1.3 |
20.6-24.2 |
62-119 |
a The animals were supplemented with 0.3 to 1 kg/d concentrate.
Farmers in India are showing increasing interest in preparing and feeding maize silage based rations to their ruminant livestock, particularly small ruminants such as growing sheep, in order to obtain optimum body weight. Maize silage was successfully supplemented, either with alfalfa hay, groundnut haulms (to meet 25% of DM requirement) or a concentrate mixture at 1.5% of body weight, in order to obtain optimum growth rate in Nellore ram lambs (Malisetty et al., 2013).
Goats
Use of maize silage in grass silage based diets
In Malaysia, the inclusion of increasing proportions of maize silage to elephant grass (Pennisetum purpureum) silage increased DM intake and growth performance of goats. The highest growth rate and OM digestibility were in goats fed 100% whole maize silage (see Table 5 below; Khaing et al., 2015).
Table 5: Effect of inclusion of maize silage in the diet (60% forage, 40% concentrate) on the performance of goats (means; SEM in brackets), adapted from Khaing et al., 2015.
Treatment (% of inclusion of maize silage
with elephant grass silage) |
OM digestibility, % |
DM intake, kg/d |
Total body weight gain, kg |
Body weight gain, g/d |
0% |
59.2 (1.43) |
277 (20) |
3.0 (0.8) |
35.7 (2.1) |
25% |
62.9 (1.43) |
339 (20) |
4.8 (0.8) |
57.1 (2.1) |
50% |
69.0 (1.43) |
381 (20) |
6.4 (0.8) |
75.6 (2.1) |
75% |
67.1 (1.43) |
407 (20) |
8.1 (0.8) |
96.0 (2.1) |
100% |
72.8 (1.43) |
464 (20) |
10.5 (0.8) |
125.4 (2.1) |
Influence of aerobic exposure of maize silage
In a preference trial, fermentation products of maize silage were affected by aerobic contamination, which had strong impacts on feed intake and preference by goats, evidenced by a decrease of DM intake after four days of exposure to oxygen. During 3h preference trials (2-way combination of maize silage and alfalfa hay), intake decreased from 651 g DM of maize silage (for 0 days of exposure) to 464 g DM of maize silage (with 8 days exposure). However, it was difficult to attribute the decline to a single cause. It was recommended that silage exposure to oxygen be limited during storage and feeding as much as possible because of its detrimental effects on intake (Gerlach et al., 2013).