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Cottonseed hulls

Description and recommendations

Common names

Cottonseed hulls, cotton seed hulls, cotton hulls


Cottonseed hulls are the outer coverings of cotton seeds, and the by-product of the dehulling step of cottonseed oil extraction. After delinting, the hulls are separated from the kernel by screening. Cottonseed hulls are a fibrous product, primarily used to feed ruminants (Hall et al., 2000). Cottonseed hulls are used for feed in bulk or pellet form. They are sometimes mixed with cottonseed meal to create a higher density product that is easier to transport and handle (Blasi et al., 2002). Cottonseed hulls are one of the best roughages used to add bulk to diets rich in protein and energy, in order to reduce digestive upsets in ruminants (Lane, 2006). Cottonseed hulls are a valuable substrate for mushroom cultivation and the spent substrate can be fed to livestock (Bae et al., 2006; Miles et al., 2004Yang et al., 1986). Cottonseed hulls have also numerous industrial uses such as plastic manufacture, oil drilling (mud additive) and furfural production (a solvent used in plastic and synthetic rubber production and in petroleum refining) (NCPA, 2011).


Cottonseed hulls are available at the mill where cottonseed oil and cottonseed meal are obtained. Due to their low density, they may be difficult to transport and are thus confined to a fairly restricted market radius (Blasi et al., 2002). They can be prepared in mixture with cottonseed meal. In the USA, the use of cottonseed hulls in pellets increased in the late 1990s (Coombs et al., 1996). The demand for hulls and the prices vary considerably with changes in the availability of other roughage feeds (NCPA, 2011).

Environmental impact

Cottonseed hulls used in complete mixed rations for dairy cows were shown to reduce greenhouse gas (CH4) production (Bo et al., 2012).

Potential constraints


Cottonseed hulls do not contain significant amounts of free gossypol: reported values are lower than 0.1% DM (Mujahid et al., 2000b; Noftsger et al., 2000), about ten times less than in the seed itself (see the Cotton seeds datasheet). Consequently, feeding animals with cottonseed hulls is less likely to result in negative effects due to gossypol. However, there has been several reports on the risk of feeding cottonseed hulls to livestock. When cottonseed hulls were fed to fattening lambs for a long period (8 months), it was found that gossypol accumulated in testis, liver, kidney, spleen and muscle (in order of decreasing content) (Tuerxhun et al., 2010). Meat inspection of 1004 Karakul sheep slaughtered in Uzbekistan in the late 1980s revealed hepatosis in 41% of the cases, cirrhosis in 11% and multiple abscesses of the liver in 4%. In a trial on 30 sheep fed 2 kg/d of cottonseed hulls, liver disease developed within 2 months and values for blood albumin and glucose were low (Norbaev, 1989). In pigs, several poisoning incidents (mortality up to 30%) have been reported in Russia in pig farms following the feeding of cottonseed meal and cottonseed hulls (Tishkov et al., 1988). 


Cottonseed hulls contain condensed tannins (3-7% DM) mainly bound to protein and fibre (Yu et al., 1996). However, it seems that the condensed tannins of cottonseed hulls do not significantly reduce the rumen protein solubility (Yu et al., 1995a; Yu et al., 1995b).

Nutritional attributes

Cottonseed hulls are a low-protein (3-9% DM), highly fibrous by-product (ADF 57-73% DM) mostly used for ruminant feeding.

Tables of chemical composition and nutritional value


Cottonseed hulls are used in ruminant feeding in areas of cotton growing. They are palatable compared to other fibrous by-products and have a stimulatory effect on feed intake in diets with limited fibre content. They were shown to increase intake compared to sunflower hulls and peanut hulls (Coppock et al., 1987).


Dairy cows

Cottonseed hulls have been included at various levels (5 to 50%) into total mixed rations or complete feeds for lactating dairy cows (Klopfenstein et al., 1981; Blasi et al., 2002). When compared to other fibrous by-products (sunflower hulls, sugarcane bagasse) or forages (Bermuda grass, maize silage, sorghum silage or sugar cane silage), cottonseed hulls generally give the same or higher milk yield and higher milk fat content (Klopfenstein et al., 1981).

For dairy cows, cottonseed hulls are a valuable fibre source when introduced into diets with high starch content because they slightly increase diet intake and nitrogen retention (Beckman et al., 2005), improve starch digestion by increasing rumination, and reduce the amount of whole maize passing through the digestive tract (Blasi et al., 2002). They tend to increase the milk fat content without modifying the milk yield (Beckman et al., 2005). However, as cottonseed hulls have a low protein content, diets that include them must be carefully balanced in crude protein (particularly when high levels of hulls are used). Cottonseed hulls can be included into complete feeds in a pelleted or non-pelleted form without any difference in milk yield (Brown et al., 1977).

The following table summarizes several experiments in the USA:

Experiment Inclusion rate Results References
Increasing levels of pelleted cottonseed hulls in place of alfalfa hay cubes 5, 15 or 25% No difference in milk yield (23-24.4 kg/d) and fat and protein content (2.9-3.2% and 2.7-2.9% respectively) Brown et al., 1977
Comparison of two levels of cottonseed hulls in TMR 30 or 40% Slight decrease of DM intake (20 kg vs 21 kg) and increase of milk fat content (3.2% vs 2.7%) at 40% level; same milk yield (22 kg/d) Olson et al., 1975
Comparison of cottonseed hulls to maize silage in TMR 30% Lower diet DM digestibility with cottonseed hulls (59 to 64%) but higher milk yield (23.4 to 20.5 kg/d) Sargent et al., 1975
Comparison of two levels of cottonseed hulls into TMR based on maize grain and soybean meal 30 or 40% No difference in DM intake (20 kg/d) or milk yield (19 kg/d); higher milk fat content with 40% cottonseed hulls (3.38 vs 3.16%) Olson et al., 1974
Beef cattle

In Brazil, cottonseed hulls could replace up to half of Pennisetum purpureum silage (60% in the diet) in a complete diet for fattening steers, increasing the daily DM intake from 6.6 kg to 8.3 kg without altering DM digestibility (54-55.6%) (Chizzotti et al., 2005).

In India, feeding complete diets based on cottonseed hulls (60%) to crossbred calves improved growth rate, feed conversion efficiency and nutrient utilization over complete diets based on wheat straw. Flaking did not improve nutrient utilization and growth but increased bulk density, which may reduce the cost of handling, transportation and storage (Ramachandran et al., 2008).

In India, crossbred bulls fed cottonseed hulls at 50% (diet DM) or a mixture of cottonseed hulls (35%) and alfafa hay (15%) supplementing a diet based on chopped sorghum straw had higher nutrient digestibilities and DM intake than bulls fed a conventional diet (concentrate + straw) (Reddy et al., 1999).

In the USA, cottonseed hulls could be used as forage source (20 or 25%) instead of alfalfa hay in a complete diet for 325 kg fattening steers (Bartle et al., 1994) or 200 kg growing beef heifers (Hale et al., 1969) without change in daily weight gain of steers and heifers (1560 g/d and 670 g/d respectively), but with a higher DM intake (+ 0.8 kg/d and + 0.52 kg/d respectively). At a higher level (30%), DM intake increased (+ 0.5 kg/d) but daily weight gain decreased to 1445 g/d (Bartle et al., 1994). Adding molasses (9%) to cottonseed hulls into the growing beef heifer diets increased growth performance to 750 g/d (Hale et al., 1969). In another comparison between cottonseed hulls and alfalfa hay where steers were fed either feed at 8% (diet DM) for 103 days, steers fed cottonseed hulls had a lower average daily gain (1930 vs 2130 g/d) and showed a tendency to have fatter carcasses and a higher (less desirable) yield grade (Markham et al., 2002).

Replacement of rice bran with up to 20% cottonseed hulls did not effect the milk yield or fat content of Murrah buffaloes (Naidu et al., 1981).



Cottonseed hulls are capable of supporting moderate growth rates in sheep. When a small amount of by-pass protein was added to the diet of cottonseed hulls + urea + 50 g lucerne + vitamins/minerals, the growth rate of lambs exceeded 130 g/day and wool growth was increased from 6 to 9 g/day. Investigation of the rumens of these animals showed that protozoa were either eliminated or in very low population densities. This could be the reason why cottonseed hulls support such reasonable growth rates, even without supplementation with by-pass protein. Intake of cottonseed hulls by sheep is higher (c. 1 kg DM/day) than would be expected of a 40% digestible feed and this is possibly associated with a rapid breakdown of the indigestible material in the rumen (Davis et al., 1989).

In Benin, the full or 50% replacement of cottonseed meal with a 60:40 mixture of cottonseed meal and cottonseed hulls to supplement Guinea grass-based diets for fattening Djallonke sheep (1 year, 18 kg) for 60 days was found to give similar performance (carcass yield and characteristics) and resulted in a net profit (Alkoiret et al., 2007).

In China, cottonseed hulls could replace the fibre source in a complete diet for fattening 3 months old sheep, resulting in the same daily weight gain and better carcass characteristics after 8 months (Tuerxhun et al., 2010)


In China, pigs were offered cottonseed hulls after use as a growth medium for 4 harvests of mushrooms (Pleurotus sp.). The optimum dose was 10% of the diet mixed with concentrates, which gave the same growth rate and carcass quality as a control diet (Yang et al., 1986).


Cottonseed hulls can be assumed to be a suitable fibre source for rabbit feeding even though no direct experiment seems to have been reported in the literature (October 2013). This assumption is based on the safe utilisation in ruminant feeding up to 20% of the diet (see Ruminants). This position is reinforced by the negligible level of gossypol observed in the cottonseed hulls even for cotton varieties with a high level on gossypol in the kernel (Yu et al., 1993). The resistance of hulls to the development of mould contamination through Aspergillus flavus and subsequent very low risk of aflatoxins development is another positive point (Mayne et al., 1966). The high level of lignin in cottonseed hulls (18-20%) should be also considered as an advantage for the control of digestive health in the rabbit (Gidenne et al., 2010).

The digestible energy content of cottonseed hulls could be estimated using equations as 5.7 MJ/kg DM and the protein digestibility as about 22% (Lebas, 2013).


Heuzé V., Tran G., Hassoun P., Lebas F., 2013. Cottonseed hulls. A programme by INRA, CIRAD, AFZ and FAO. Last updated on December 15, 2013, 20:10


Tables of chemical composition and nutritional value

Main analysis Unit Avg SD Min Max Nb  
Dry matter % as fed 90.6 1.8 87.4 93.3 18  
Crude protein % DM 5.1 1.6 2.6 8.7 35  
Crude fibre % DM 53.0 7.5 35.0 64.8 21  
NDF % DM 85.3 5.8 71.6 91.6 18  
ADF % DM 63.0 4.9 56.5 73.4 17  
Lignin % DM 19.1 3.2 11.5 23.2 15  
Ether extract % DM 3.2 1.9 1.0 8.1 16  
Ash % DM 2.9 0.4 2.5 3.7 29  
Starch (polarimetry) % DM 4.2       1  
Gross energy MJ/kg DM 19.8 0.4 18.3 19.8 3 *
Minerals Unit Avg SD Min Max Nb  
Calcium g/kg DM 1.3 0.4 1.0 2.2 19  
Phosphorus g/kg DM 1.0 0.5 0.7 2.9 20  
Potassium g/kg DM 7.1 3.0 4.5 12.4 15  
Sodium g/kg DM 0.2 0.0 0.2 0.2 10  
Magnesium g/kg DM 1.3 0.5 0.6 2.0 17  
Manganese mg/kg DM 22 4 19 33 10  
Zinc mg/kg DM 4 5 2 20 11  
Copper mg/kg DM 69 22 7 95 11  
Iron mg/kg DM 172 46 139 308 11  
Amino acids Unit Avg SD Min Max Nb  
Alanine % protein 3.7       1  
Arginine % protein 2.9       1  
Aspartic acid % protein 19.7       1  
Glutamic acid % protein 8.3       1  
Glycine % protein 2.9       1  
Histidine % protein 1.4       1  
Isoleucine % protein 2.6       1  
Leucine % protein 4.0       1  
Lysine % protein 3.1       1  
Phenylalanine % protein 2.6       1  
Proline % protein 3.4       1  
Serine % protein 4.6       1  
Threonine % protein 2.6       1  
Tyrosine % protein 4.6       1  
Valine % protein 3.1       1  
Secondary metabolites Unit Avg SD Min Max Nb  
Tannins, condensed (eq. catechin) g/kg DM 51.5   51.0 52.0 2  
Gossypol, free mg/kg DM 430 191 160 680 12  
Ruminant nutritive values Unit Avg SD Min Max Nb  
OM digestibility, ruminants % 42.9 7.2 36.9 50.9 3  
Energy digestibility, ruminants % 41.1         *
DE ruminants MJ/kg DM 8.2         *
ME ruminants MJ/kg DM 6.5         *
ME ruminants (gas production) MJ/kg DM 6.6   6.1 7.1 2  
ME ruminants (FAO, 1982) MJ/kg DM 7.3       1  
Nitrogen digestibility, ruminants % 2.6 27.6 -30.9 36.4 4  
a (N) % 3.0   3.0 3.0 2  
b (N) % 40.5   37.0 44.0 2  
c (N) h-1 0.030   0.021 0.038 2  
Nitrogen degradability (effective, k=4%) % 20   16 24 2 *
Nitrogen degradability (effective, k=6%) % 16   13 20 2 *
Rabbit nutritive values Unit Avg SD Min Max Nb  
DE rabbit MJ/kg DM 5.7         *
Nitrogen digestibility, rabbit % 22.0         *

The asterisk * indicates that the average value was obtained by an equation.


AFZ, 2011; Alibes et al., 1990; Ashes et al., 1978; Beckman et al., 2005; Bhatti et al., 1995; Bo et al., 2012; Brown et al., 1977; Chizzotti et al., 2005; CIRAD, 1991; CIRAD, 2008; Garleb et al., 1988; Gartner et al., 1975; Göhl, 1970; Gowda et al., 2004; Hale et al., 1969; Henderson et al., 1984; Hsu et al., 1987; Krishna, 1985; Lirette et al., 1993; Mujahid et al., 2000; Neumark, 1970; Noftsger et al., 2000; Richard et al., 1989; Wheeler et al., 1979; Yu et al., 1995; Yu et al., 1995

Last updated on 19/11/2013 16:58:38



Alkoiret, T. I. ; Manne, A. A. S. ; Gbangboche, A. B. ; Attakpa, E. Y., 2007. Fattening performance of Djallonke sheep supplemented with cotton seed husks in Benin. Livest. Res. Rural Dev., 19 (10): 141 web icon
Arshami, J. ; Ruttle, J. L., 1988. Effects of diets containing gossypol on spermatogenic tissues of young bulls. Theriogenology, 30 (3): 507-516 web icon
Bae, J. S. ; Kim, Y. I. ; Jung, S. H. ; Oh, Y. G. ; Kwak, W. S., 2006. Evaluation on feed-nutritional value of spent mushroom (Pleurotus ostreatus, Pleurotus eryngii, Flammulina velutipes) substrates as a roughage source for ruminants. J. Anim. Sci. Technol., 48 (2): 237-246 web icon
Bartle, S. J.; Preston, R. L.; Miller, M. F., 1994. Dietary energy source and density: effects if roughage source, roughage equivalent, tallow level and steer type on feedlot performance and carcass characteristics. J. Anim. Sci., 72 (8): 1943-1953 web icon
Beckman, J. L.; Weiss, W. P., 2005. Nutrient digestibility of diets with different fiber to starch ratios when fed to lactating dairy cows. J. Dairy Sci., 88 (3): 1015-1023 web icon
Blasi, D. A. ; Drouillard, J., 2002. Composition and feeding value of cottonseed feeds products for beef cattle. Kansas State University, May 2002. 22p web icon
Bo, Y. K.; Yang, H. J.; Wang, W. X.; Liu, H.; Wang G. Q.; Yu, X., 2012.  Metabolisable energy, in situ rumen degradation and in vitro fermentation characteristics of linted cottonseed hulls, delinted cottonseed hulls and cottonseed linter residue. Asian-Aust. J. Anim. Sci., 25 (2): 240-247 web icon
Brown, W. H. ; Whiting, F. M. ; Daboll, B. S. ; Turner, R. J. ; Schuh, J. D., 1977. Pelleted and non-pelleted cottonseed hulls for lactating dairy cows. J. Dairy Sci., 62 (6): 919-923 web icon
Chizzotti, M. L.; Valadares Filho, S. de C.; Leao, M. I. ; Valadares, R. F. D.; Chizzotti, F. H. M.; Magalhaes, K. A.; Marcondes, M. I., 2005. Partial replacement of elephant grass silage with cottonseed hulls. 1. Intake, degradability, and apparent ruminal, intestinal, and total tract digestibilities in steers. Rev. Bras. Zootec., 34 (6): 2093-2102 web icon
Coombs, D. F. ; Pontif, J. E., 1996. An evaluation of cottonseed hulls and cottonseed hull pellets in beef cattle diets. Annual Research Sumamary, Dean Lee Research Station, LAES, p. 3.
Coppock, C. E. ; Lanham, J. K. ; Horner, J. L., 1987. A review of the nutritive value and utilization of whole cottonseed, cottonseed meal and associated by-products by dairy cattle. Anim. Feed Sci. Technol., 18 (2): 89-129 web icon
Davis, J. J. ; Leng, R. A., 1989. Cottonseed hulls as a feed for sheep. Recent Advances in Animal Nutrition in Australia 1989. FARRELL, D.J. (Ed.). University of New England, Armidale. 33A web icon
Erdman, R. A. ; Proctor, G. H. ; Vandersall, J. H., 1986. Effect of rumen ammonia concentration on in situ rate and extent of digestion of feedstuffs. J. Dairy Sci., 69 (9): 2312-2320 web icon
Garleb, K. A. ; Fahey, G. C. Jr. ; Lewis, S. M. ; Kerley, M. S. ; Montgomery, L., 1988. Chemical composition and digestibility of fiber fractions of certain by-product feedstuffs fed to ruminants. J. Anim. Sci., 66 (10): 2650-2662 web icon
Gidenne, T.; García, J.; Lebas, F.; Licois, D., 2010. Nutrition and feeding strategy: interactions with pathology. In: Nutrition of the rabbit - 2nd edition. de Blas, C.; Wiseman, J. (Eds). CAB International, UK web icon
Hale, W. H.; Lambeth, C.; Theurer, B.; Ray, D. E., 1969. Digestibility and utilization of cottonseed hulls by cattle. J. Anim. Sci., 29 (5): 773-776 web icon
Hall, M. B.; Akinyode, A., 2000. Cottonseed hulls: working with a novel fiber source. In: Proc. 11th Ann. Florida Rumin. Gainesville, FL: Nutr. Symp.: 179–186 web icon
Hsu, J. T. ; Faulkner, D. B. ; Garleb, K. A. ; Barclay, R. A. ; Fahey, G. C. ; Berger, L. L., 1987. Evaluation of corn fiber, cottonseed hulls, oat hulls and soybean hulls as roughage sources for ruminants. J. Anim. Sci., 65 (1): 244-255 web icon
Ikitoo, E.C., 2011. Gossypium hirsutum L.. Record from Protabase. Brink, M. & Belay, G. (Editors). PROTA (Plant Resources of Tropical Africa / Ressources végétales de l’Afrique tropicale), Wageningen, Netherlands web icon
Klopfenstein, T.; Owen, F. G., 1981. Value and potential use of crop residues and by-products in dairy rations. J. Dairy Sci., 64 (6): 1250-1268 web icon
Kononoff, P. J.; Heinrichs, A. J., 2002. The effect of corn silage particle size and cottonseed hulls on cows in early lactation. J. Dairy Sci., 86 (7): 2438-2451 web icon
Lane, C. D. Jr., 2006. Feeding cottonseed and cotton byproducts to beef cattle. Univ. Tennessee. The University of Tennessee Extension. AS - B 352 web icon
Lebas, F., 2013. Estimation de la digestibilité des protéines et de la teneur en énergie digestible des matières premières pour le lapin, avec un système d’équations. 15ème Journées de la Recherche Cunicole, Le Mans, 19-20 novembre 2013
Markham, C. E. ; Krehbiel, C. R. ; Gill, D. R. ; Lalman, D. L. ; McBeth, L. J. ; Ball, R. L., 2002. Effects of fiber source on feedlot performance and carcass characteristics of yearling steers. Animal Science Research Report - Agricultural Experiment Station, Oklahoma State University, P-993 web icon
Mayne, R. Y.; Pons Jr, W. A.; Franz Jr, A. O.; Goldblatt, L. A., 1966. Elaboration of aflatoxin on cottonseed products by Aspergillus flavus. J. Am. Oil Chem. Soc., 43: 251-253 web icon
Miles, P. G.; Shu-Ting Chang, 2004. Mushrooms: cultivation, nutritional value, medicinal effect, and environmental impact. 2nd Edition. CRC Press, 2004 web icon
Mujahid, A. ; Abdullah, M. ; Barque, A. R. ; Gilani, A. H., 2000. Nutritional value of cottonseeds and its derived products : I. Physical fractionations and proximate composition. Asian-Aust. J. Anim. Sci., 13 (3): 348-355 web icon
Mujahid, A. ; Abdullah, M. ; Barque, A. R. ; Gilani, A. H., 2000. Nutritional value of cottonseeds and its derived products : II. Free gossypol, available lysine and in vitro protein digestibility. Asian-Aust. J. Anim. Sci., 13 (3): 356-359 web icon
Naidu, M. M. ; Das, C. T. ; Reddy, M. R., 1981. Effect of incorporating cottonseed hulls in concentrates on the quantity and quality of milk in dairy buffaloes. Agricultural Science Digest, India, 1 (4): 265-269
NCPA, 2011. Petition for renewable fuel pathway for biodiesel using cottonseed oil - Renewable Fuel Standard 40 C.F.R. §80.1416. National Cottonseed Products Association, Tennessee, USA web icon
NCPA, 2012. Cottonseed feed products guide. National Cotton Products Association web icon
Noftsger, S. M.; Hopkins, B. A.; Diaz, D. E.; Brownie, C.; Whitlow, L. W., 2000. Effect of whole and expanded-expelled cottonseed on milk yield and blood gossypol. J. Dairy Sci., 83 (11): 2539–2547 web icon
Norbaev, K. N., 1989. Aetiology and semiology of hepatosis in fattening sheep. Veterinariya Moskva, 8: 57-59
Olson, H. H.; Santos, K. A. S.; McCoy, G. C., 1974. Percent roughage in corn and milo complete feeds. J. Dairy Sci., 57 (10): 1260-1262 web icon
Olson, H. H.; McCoy, G. C., 1975. Complete feeds with 30 and 40 percent cottonseed hull for lactating dairy cows. J. Dairy Sci., 58 (5): 747 (abstract) web icon
Ramachandran, M. ; Singhal, K. K., 2008. Cottonseed hulls based complete diets for crossbred calves. Livest. Res. Rural Dev., 20 (11): 176 web icon
Reddy, G. V. N. ; Reddy, M. R., 1999. Nutrient utilization and rumen fermentation pattern of cotton seed hulls based complete diets in crossbred bulls. Indian J. Anim. Nutr., 16 (1): 6-11
Sargent, A. L.; McCoy, G. C.; Olson, H. H., 1975. Digestibility relationships between complete feeds with corn silage and cottonseed hulls. J. Dairy Sci., 58 (5): 762 (abstract) web icon
Tishkov, A. I., 1988. Post mortem diagnosis of gossypol poisoning in pigs. Veterinariya, Moscow, 11: 58-59
Tuerxhun PaXia ; Wen ZhengShun ; Pan XiaoLiang ; Shen SiJun ; Xu ZiRong, 2010. Effect of cottonseed hull on growth performance and tissue residues of gossypol in fine-wool male sheep. Zhejiang University (Agricult. Life Sci., 36 (3): 306-310 web icon
Yang, S. Y. ; Huang, Y. B. ; Du, P. Z. ; Chu, X. S. ; Meng, M. Z., 1986. Feeding mushroom growth medium to fattening pigs. Animal Husbandry and Veterinary Medicine, China 18 (5): 202-203
Yu, F. ; Barry, T. N. ; Moughan, P. J. ; Wilson, G. F., 1993. Condensed tannin and gossypol concentrations in cottonseed and in processed cottonseed meal. J. Sci. Food Agric., 63 (1): 7-15 web icon
Yu, F. ; McNabb, W. C. ; Barry, T. N. ; Waghorn, G. C., 1995. Effect of condensed tannin in cottonseed hulls upon the in vitro degradation of cottonseed kernel proteins by rumen microorganisms. J. Sci. Food Agric., 69 (2): 223-234 web icon
Yu, F. ; Barry, T. N. ; McNabb, W. C. ; Moughan, P. J. ; Wilson, G. F., 1995. Effect of bound condensed tannin from cottonseed upon in situ protein solubility and dry matter digestion in the rumen. J. Sci. Food Agric., 69 (3): 311–319 web icon
Yu, F. ; Moughan, P. J. ; Barry, T. N., 1996. The effect of cottonseed condensed tannins on the ileal digestibility of amino acids in casein and cottonseed kernel. Br. J. Nutr., 75 (5): 683-698 web icon

Image credits

Image credits

Picture title Credits License
Cottonseed hulls Ella Liliane Mutuyimana / Kigali Farms, Rwanda CC BY 3.0
Cottonseed hulls Laurent Demuynck / Kigali Farms, Rwanda CC BY 3.0
Bags of cottonseed hulls Laurent Demuynck / Kigali Farms, Rwanda CC BY 3.0
Cottonseed hulls for mushroom production Laurent Demuynck / Kigali Farms, Rwanda CC BY 3.0